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Catalogue of the Living Bivalvia of the Eastern Pacific Ocean ...

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32. Chlamys islandica (Müller 1776) according to MacNeil (1967) does<br />

not occur in <strong>the</strong> <strong>Pacific</strong> <strong>Ocean</strong> or Bering Sea, though Grau (1959) considered<br />

C. behringiana (Middendorff 1849) and C. albida (Arnold 1906) to be<br />

subspecies <strong>of</strong> C. islandica. I believe <strong>the</strong>m to be distinct but on <strong>the</strong> o<strong>the</strong>r<br />

hand, species proposed by MacNeil, appear tome to be mere variants <strong>of</strong> C.<br />

islandica.<br />

33. Hinnites (DeFrance 1821) is usually accorded full generic status. I am<br />

unable to find distinguishing characters and do not consider <strong>the</strong> late cementation<br />

and idiomorphic growth sufficient to accord more than subgeneric<br />

separation.<br />

34. Semipallium zeteki (Hertlein 1935) was proposed as a replacement<br />

name for <strong>the</strong> preoccupied Pecten digitatus (Hinds 1844), but nothing corresponding<br />

to <strong>the</strong> description is known from <strong>the</strong> eastern <strong>Pacific</strong>. The taxon is<br />

probably referable to S. vexillum (Reeve 1853) from <strong>the</strong> western <strong>Pacific</strong>.<br />

35. Cyclopecten imbrifer (Lovén 1847) probably does not occur in <strong>the</strong><br />

eastern <strong>Pacific</strong>, but is included on <strong>the</strong> basis <strong>of</strong> material from Oregon that may<br />

be a new species, but <strong>the</strong> relationship to C. benthalis (Grau 1959) must be<br />

resolved.<br />

36. Cyclopecten incotigruus (Dall 1916) is known from <strong>the</strong> holotype only;<br />

though <strong>the</strong> type locality is cited as San Diego, California, <strong>the</strong> specimen label<br />

states Cedros Island, Baja California.<br />

37. Cyclopecten vitreus (Gmelin 1791) seems to be <strong>the</strong> species with <strong>the</strong><br />

widest distribution, but may include a complex <strong>of</strong> species that may be<br />

separable with <strong>the</strong> study <strong>of</strong> shell ultrastructure.<br />

38. Pecten berlyi nom. nov. is proposed for <strong>the</strong> preoccupied P. lunaris<br />

(Berry 1963).<br />

39. Spondylus tenebrosus (Reeve 1856) has been recorded from Clipperton<br />

Island (Salvat & Ehrhardt 1970).<br />

40. Spondylus victoriae (Sowerby 1859) is widespread throughout <strong>the</strong><br />

Gulf <strong>of</strong> California and is well represented in <strong>the</strong> collection <strong>of</strong> <strong>the</strong> Allan<br />

Hancock Foundation. Though overlooked by modern workers, <strong>the</strong> name<br />

was used for Recent and Pliocene species by Durham (1950).<br />

41. Anomia chinensis (Philippi 1849) was incidentally introduced from<br />

Japan to Willapa Bay, Washington, but did not become established.<br />

42. Pododesmus cepio (Gray 1850) is frequently merged with P. macrochisma<br />

(Deshayes 1839) which is confined to Alaska.<br />

43. Lucina approximata (Dall 1901) probably is represented by a complex<br />

<strong>of</strong> several species.<br />

44. Lucina mazatlanica (Carpenter 1857) requires careful evaluation. It<br />

has been reported from <strong>the</strong> Gulf <strong>of</strong> California in 1043 m (Emerson and Puffer<br />

1957) ,<br />

45. Thyasirinae subfamily nov. is proposed for members <strong>of</strong> <strong>the</strong> family<br />

Thyasiridae that are deeply infaunal, usually in regions <strong>of</strong> low productivity<br />

and low oxygen tension. The unsculptured shell has sharp dorsal folds<br />

accompanied by one or two radial sulci. The most pr<strong>of</strong>ound modifications<br />

are in <strong>the</strong> s<strong>of</strong>t parts, particularly <strong>the</strong> long vermiform foot, with terminal<br />

secretory bulb, which forms <strong>the</strong> mucoid inhalant tube. The anterior adductor<br />

muscle is elongated by <strong>the</strong> rotation <strong>of</strong> <strong>the</strong> body relative to <strong>the</strong> shell. The type<br />

genus is Thyasira Leach in Lamarck 1818. Included genera are Conchocele<br />

(Gabb 1866), Maorithyas (Fleming 1950), Parathyasira (Iredale 1930) and<br />

Philis (Fischer 1861).<br />

46. Conchocele excavata (Dall 1901) is usually placed in Thyasira sensu<br />

stricto, but <strong>the</strong> shell and anatomy show that it should be assigned to<br />

Conchocele. Shallow water (18-90 m) records from <strong>the</strong> Gulf <strong>of</strong> California<br />

(Parker 1964) are not this species.<br />

47. Axinopsidinae subfamily nov. is proposed for members <strong>of</strong> <strong>the</strong> family<br />

Thyasiridae that are members <strong>of</strong> <strong>the</strong> shallow infauna. Unlike <strong>the</strong> Thyasirininae,<br />

<strong>the</strong> group lacks shell folds or sulci, <strong>the</strong> hinge is streng<strong>the</strong>ned by<br />

tubercles or pseudodentition, and <strong>the</strong> lunule is well developed. The subfamilies<br />

have arborescent digestive diverticula, but <strong>the</strong> foot is much shorter<br />

68<br />

in <strong>the</strong> Axinopsininae and lacks <strong>the</strong> hypertrophied terminal portion. I consider<br />

<strong>the</strong> proposed subfamily, dating from <strong>the</strong> Pliocene, only distantly related to<br />

<strong>the</strong> Thyasirinae which arose in <strong>the</strong> Cretaceous. The type genus is Axinopsida<br />

Keen and Chavan in Chavan 1951. Included genera are Adontorhina (Berry<br />

1947) and Axinulus (Verrill & Bush 1898).<br />

48. Felaniella parilis (Conrad 1848), described as a Miocene fossil, was<br />

first united with living material by Grant & Gale (1931) though largely<br />

ignored by subsequent workers, review <strong>of</strong> a large suite <strong>of</strong> living and fossil<br />

material from California and Washington (Pliocene), confirms my opinion.<br />

49. Kelliinae subfamily nov. is proposed to contain members <strong>of</strong> <strong>the</strong> family<br />

Kelliidae with strong dentition, mantle folds not extendable over <strong>the</strong> shell<br />

surface and <strong>the</strong> foot not modified into a wide creeping sole. The type genus is<br />

Kellia (Turton 1822). Included genera are Aligena (Lea 1846), Diplodontina<br />

(Stempell 1899), and Odontogena Cowan 1964.<br />

50. Aligena pisum (Dall 1908) is known only from a left valve, eroded and<br />

immature. It has been assigned to Axinulus by Harry (1969), but certainly is<br />

not a member <strong>of</strong> that genus. This taxon may best be considered a nomen<br />

dubium.<br />

51. Borniinae subfamily nov. is proposed to contain <strong>the</strong> genera Bornia<br />

(Philippi 1836), Rhamphidonta (Bernard 1975) and Solecardia (Conrad<br />

1849). Unlike o<strong>the</strong>r members <strong>of</strong> <strong>the</strong> family Kelliidae, this group has <strong>the</strong><br />

hinge comparatively weakly developed, <strong>the</strong> shell thin and elongated. The<br />

mantle inner fold is capable <strong>of</strong> great extension to cover <strong>the</strong> entire shell<br />

valves, and <strong>the</strong> foot modified into a wide creeping disc. The type genus is<br />

Bornia (Philippi 1836).<br />

52. Ercyiniinae subfamily nom, trans!, is proposed for member genera <strong>of</strong><br />

<strong>the</strong> family Lasaeidae with elongated compressed shells, weak to moderate<br />

hinge plate, laterals lamelliform, ligament extending backwards. The type<br />

genus is Erycina Lamarck 1805. Also included is Amerycina Chavan 1959.<br />

53. Lasaeinae subfamily nom, trans!, is proposed for <strong>the</strong> distinctive genus<br />

Lasaea (Brown 1827) <strong>of</strong> <strong>the</strong> family Lasaeidae. The subfamily is characterized<br />

by a rounded minute shell, nearly vertical beaks, and a thick, heavy<br />

hinge with a protruding cardinal in <strong>the</strong> left valve. The lateral teeth are similar<br />

in both valves and continuous with o<strong>the</strong>r teeth.<br />

54. Montacutinae subfamily nom, trans!, is proposed for genera <strong>of</strong> <strong>the</strong><br />

family Montacutidae with subequilateral shells, thickened lateral teeth and<br />

minute resilium. The type genus is Montacuta (Turton 1822). The o<strong>the</strong>r<br />

included genus is Montacutona (Yamamoto and Habe 1959).<br />

55. Mysellinae subfamily nov. is proposed for <strong>the</strong> very distinct genus<br />

Mysella (Angas 1877) which should probably be divided into several<br />

genera. All species have a large resilifer, frequently bounded by raised<br />

margins or teeth forming a chondrophore-like structure.<br />

56. Orobitellinae subfamily nov. is proposed for a number <strong>of</strong> genera <strong>of</strong> <strong>the</strong><br />

family Montacutidae, with usually inequilateral shells and oblique poorly<br />

developed resilifers. The right valve displays a strong anterior lateral tooth.<br />

The type genus is Orobitella (Dall 1900). Included genera are lsorobitella<br />

(Keen 1962), Neaeromya (Gabb 1873), Pythinella (Dall 1899), and tentatively,<br />

Scioberetia (Bernard 1895).<br />

57.Thecodontinae subfamily nov. is proposed for very inequilateral genera<br />

<strong>of</strong> <strong>the</strong> Montacutidae, with projecting prosogyrate beaks, resilifer oblique.<br />

Left valve with anterior lateral tooth slightly serrated and curved, following<br />

shell margin. The type genus is Thecodonta (A.Adams 1864). The east<br />

<strong>Pacific</strong> representative is Pristes (Carpenter 1864).<br />

58. Basterotia quadrata (Hinds 1843), described without locality.<br />

According to Olsson (1961) it was cited by Dall as a member <strong>of</strong> <strong>the</strong> east<br />

<strong>Pacific</strong> fauna. I am unable to find mention <strong>of</strong> any such species by Dall, but<br />

Paetel (1890) did include it in his catalogue. The species is a common warm<br />

water west Atlantic <strong>Ocean</strong> and Caribbean species.<br />

59. Cyclocardia gouldii (Dall 1903) was described from 1503m <strong>of</strong>f San<br />

Diego, California, but no o<strong>the</strong>r material from this well-collected area<br />

matches <strong>the</strong> type material. I consider this to be a misplaced Atlantic C.<br />

borealis (Conrad 1831), <strong>the</strong> elongate form sometimes accorded subspecific<br />

status as C. borealis novangliae (Morse 1869). This conclusion may be

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