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A Classification of the Recent Elasmobranchii

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FAMILY RHINOBATIDAE<br />

Genus RHINOBATOS Linck 1790<br />

Rhinobatos ARISTOTLE<br />

Rhinobates JOVIUS 1524<br />

Rhinobatos WOTTON 1552<br />

Rhinobatus RONDELET 1554<br />

Rhinobatus sive squatinoraia FABIUS COLUMNA 1592<br />

Rhinobatus KLEIN 1742; 1775 (Inadmissible work) type, Raja rhinobatos LINN. 1758 (assumed<br />

tautotype)<br />

Leiobatus KLEIN 1742; 1775 (Inadmissible work)<br />

Rhinobatos LINCK 1790 Raja rhinobatos LINN. 1758 (assumed tautotype)<br />

Rhinobatus WALBAUM 1792 on KLEIN 1775 (Inadmissible work) type, Raja rhinobatos LINN.<br />

1758 (assumed tautotype)<br />

Leiobatus WALBAUM 1792 on KLEIN 1775 ( Inadmissible work)<br />

Rhinobatus BLOCH & SCHNEIDER 1801 type, Raja rhinobatos LINN. 1758<br />

Leiobatus RAFINESQUE 1810 type, Leiobatus panduratus RAFINESQUE 1810<br />

( = R.. rhinobatos), not precluded by KLEIN and WALBAUM. Used as a subgenus by Fowler 1941,<br />

following Norman 1926, but no type designated. If panduratus is rhinobatos this cannot be used as<br />

Rafinesque 1810. is fully available.<br />

Squatinoraja NARDO 1824, type Squatinoraja colonna NARDO 1824 (= Rhinobatos columnae<br />

BONAPARTE = Raja rhinobatos LINN. )<br />

Rhinobatis BLAINVILLE 1825, type Raja rhinobatos LINN. 1758<br />

Leiobatis BLAINVILLE 1825, type Leiobatus panduratus RAF. 1810 ( = Raja rhinobatos LINN.<br />

1758)<br />

Rhinobates SWAINSON 1839 type Raja rhinobatos LINN. 1758<br />

Rhinobatus (Syrrhina) MUELLER & HENLE 1841 (part) type, Rhinobatus columnae<br />

BONAPARTE 1836 (= rhinobatos) by subsequent designation <strong>of</strong> J.R. Norman 1966 (Manuscript).<br />

Mueller & Henle also included blochii, annulatus (in Rhinobatos), brevirostris (now Zapteryx),<br />

banksii and bougainvillii (now Aptychotrema).<br />

Glaucostegus BONAPARTE 1846 type Rhinobatus cemiculus ST. HILAIRE 1817<br />

Squatiniraja AGASSIZ 1846 ( emendation <strong>of</strong> NARDO )<br />

Rhinobatus (Acroteriobatis) GILTAY 1928, type Rhinobatus (Syrrhina) annulatus M.&H. 1841<br />

Subgenus ACROTERIOBATIS<br />

Rhinobatos (Scobatus) WHITLEY 1939, type Rhinobatus granulatus BLAINV. 1816.<br />

Rhinobatos (Platypornax) WHITLEY 1939, type Rhinobatus (Rhinobatus) thouini M.& H. 1841.<br />

Group I (Subgenus RHINOBATUS)<br />

A1a rhinobatos<br />

A1b hynnicephalus<br />

A2a schlegeli<br />

A2b holcorhynchus<br />

Group 2 (Subgenus ACROTERIOBATIS)<br />

A3a annulatus<br />

A3b leucospilus<br />

A3c zanzibarensis<br />

A4 blochii<br />

Group 3<br />

B1a halavi<br />

B1b cemiculus<br />

B2a typus<br />

B2b microphthalmus<br />

B3 obtusus<br />

B4a percellens<br />

B4b horkelii<br />

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