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The Isopods of Abyssal Depths in the Atlantic Ocean

The Isopods of Abyssal Depths in the Atlantic Ocean

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Type locality: North <strong>Atlantic</strong>, L.G.O. Biotrawl No.<br />

7, type only, cat. no. 1-191.<br />

ABYSSAL ATLANTIC ISOPODS 183<br />

Distribution: Known only from type locality.<br />

Aff<strong>in</strong>ities: Unique.<br />

Family: ECHINOTHAMBEMIDAE Menzies<br />

Synonyms: Ech<strong>in</strong>othambemvdae Menzies, 1956a, pp.<br />

9-10.<br />

Diagnosis: Paraselloidea with fused or free head,<br />

eyes lack<strong>in</strong>g. Mandibles normal, molar process well<br />

developed and expanded a truncated apex. Antennae<br />

shorter than body, about twice <strong>the</strong> length <strong>of</strong> cephalon.<br />

All peraeopods simple, 2-7 simple walk<strong>in</strong>g legs; dactyl<br />

with two term<strong>in</strong>al claws. Uropoda term<strong>in</strong>al with<br />

peduncle. Last one or two peraeonal somites fused<br />

with pleon. First three articles <strong>of</strong> maxillipedal palp<br />

expanded, as wide as endite. Coxal plates lack<strong>in</strong>g.<br />

First peduncular article <strong>of</strong> first antenna much<br />

expanded. Anus conta<strong>in</strong>ed with<strong>in</strong> <strong>the</strong> branchial<br />

chamber.<br />

Composition: Formerly monotypic (Menzies, 1956a,<br />

p. 10). A new genus is added here and <strong>the</strong> diagnosis<br />

has accord<strong>in</strong>gly been altered to allow its <strong>in</strong>clusion.<br />

Genus: ECHINOTHAMBEMA Menzies<br />

Type species: Ech<strong>in</strong>othambema ophiuroides Menzies<br />

1956a, p. 11.<br />

Synonyms: Ech<strong>in</strong>othambema Menzies, 1956a, pp.<br />

10-11.<br />

Diagnosis: Same as for family. Mandible without<br />

palp.<br />

Composition: One abyssal <strong>Atlantic</strong> species.<br />

Ech<strong>in</strong>othambema ophiuroides Menzies<br />

Figure 62 N-O<br />

Synonyms: Ech<strong>in</strong>othambema ophiuroides Menzies,<br />

1956a, p. 11.<br />

Diagnosis: Ech<strong>in</strong>othambema with first antenna with<br />

six articles, last one-third longer than penultimate<br />

article. Lateral borders <strong>of</strong> peraeonal somites each<br />

with expanded and pronounced marg<strong>in</strong> bear<strong>in</strong>g a<br />

s<strong>in</strong>gle stout seta. Body sharply granulate. Maxilliped<br />

with two coupl<strong>in</strong>g hooks. Cephalon at antero-lateral<br />

Diagnosis: Paraselloidea with cephalon free, eyes<br />

lack<strong>in</strong>g. Mandibles normal, molar process well<br />

developed and expanded at truncated apex. Antennae<br />

shorter than body. All peraeopods simple walk<strong>in</strong>g<br />

legs, dactyls with two term<strong>in</strong>al claws. Uropod absent.<br />

Pleon with one somite only. All peraeonal somites<br />

dist<strong>in</strong>ct. First three articles <strong>of</strong> maxillipedal palp as<br />

wide as endite. Coxal plates lack<strong>in</strong>g.<br />

marg<strong>in</strong> with a pair <strong>of</strong> swell<strong>in</strong>gs, each bear<strong>in</strong>g a stout<br />

seta. Mandibles without a palp.<br />

Measurements: Holotype ambisexual 5.00 mm.<br />

long, 1.5 mm. wide.<br />

Type locality: North <strong>Atlantic</strong>, L.G.O. Biotrawl<br />

No. 1, type only, cat. no. 11760 A.M.N.H.<br />

Distribution: Known only from type locality.<br />

Aff<strong>in</strong>ities: Unique.<br />

VEMATHAMBEMA, new genus<br />

Type species: Vemathambema elongata, new species.<br />

Synonyms: None.<br />

Diagnosis: Ech<strong>in</strong>othambemidae with cephalon<br />

fused to first peraeonal somite and last somite <strong>of</strong><br />

peraeon fused to pleon. First article <strong>of</strong> first antenna<br />

not expanded but stout and elongated as long as<br />

second.<br />

Composition: Monotypic abyssal.<br />

Aff<strong>in</strong>ities: Related to Ech<strong>in</strong>othambema <strong>in</strong> <strong>the</strong> absence<br />

<strong>of</strong> coxal plates and <strong>the</strong> fusion <strong>of</strong> peraeonal<br />

somite 7 with <strong>the</strong> pleon and <strong>in</strong> <strong>the</strong> absence <strong>of</strong> a<br />

mandibular palp. Uropoda (miss<strong>in</strong>g) probably with<br />

peduncle.<br />

Vemathambema elongata, new species<br />

Figure 62 J-M<br />

Synonyms: None.<br />

Diagnosis: Vemathambema with smooth pleon, apex<br />

rounded, body without sp<strong>in</strong>es or setae. Frons <strong>of</strong><br />

cephalon concave, devoid <strong>of</strong> sp<strong>in</strong>es or setae<br />

Measurements: Holotype female length 5.2 mm<br />

width pleon 0.9 mm.<br />

Type locality: South <strong>Atlantic</strong>, L.G.O. Biotrawl<br />

No. 15, type only, cat. no. 1-112.<br />

Distribution: Known only from type locality.<br />

Aff<strong>in</strong>ities: Unique.<br />

Family: THAMBEMATIDAE<br />

Composition: Monotypic. <strong>The</strong> family resembles<br />

<strong>the</strong> Jaeropsidae <strong>in</strong> many significant respects,<br />

notably <strong>in</strong> <strong>the</strong> details <strong>of</strong> <strong>the</strong> structure <strong>of</strong> <strong>the</strong><br />

maxillipeds and <strong>the</strong> shape <strong>of</strong> <strong>the</strong> front <strong>of</strong> <strong>the</strong><br />

head and <strong>the</strong> antennae structure. <strong>The</strong> mandibular<br />

structure, however, clearly <strong>in</strong>dicates its dist<strong>in</strong>ctiveness.

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