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The Isopods of Abyssal Depths in the Atlantic Ocean

The Isopods of Abyssal Depths in the Atlantic Ocean

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90<br />

ROBERT J. MRNZIES<br />

10. Head subcircular, mandibles small and not<br />

extend<strong>in</strong>g to marg<strong>in</strong> <strong>of</strong> frons . . . serrata Richardson<br />

(709 meters, Albatross, N. <strong>Atlantic</strong>, south <strong>of</strong> Martha's<br />

V<strong>in</strong>eyard)<br />

10. Head quadrate, mandibles<br />

normal tuberculata Richardson<br />

(1132 meters, Albatross, 37° 22'.30" N., 137° 47' E.,<br />

<strong>of</strong>f Japan)<br />

11. Peraeopods strongly sp<strong>in</strong>ous, head small<br />

(much narrower than peraeon) . . stygia (G. O. Sars)<br />

(535-2391 meters, N. <strong>Atlantic</strong>)<br />

11. Peraeopods weakly sp<strong>in</strong>ous, head large (nearly as<br />

wide as peraeon) 12<br />

12. Somite 4 <strong>of</strong> peraeon with a sulcus on<br />

dorsal suface at midl<strong>in</strong>e caeca Richardson<br />

(2638 meters, N. <strong>Atlantic</strong>)<br />

12. Somite 4 <strong>of</strong> peraeon without a sulcus on dorsal<br />

surface at midl<strong>in</strong>e 13<br />

13. Lateral border <strong>of</strong> head strongly convex.<br />

Lateral marg<strong>in</strong> <strong>of</strong> peraeon convex . . bicolor Hansen<br />

(1537 meters, N. <strong>Atlantic</strong>)<br />

13. Lateral border <strong>of</strong> head scarcely convex, borders<br />

<strong>of</strong> peraeon subparallel albescens Hansen<br />

(842-1018 meters, N. <strong>Atlantic</strong>)<br />

Genus: GNATHIA Leach<br />

Gnathia vemae, new species<br />

Figure 2<br />

Figure 2. Gnathia vemae, n. sp. A: male holotype;<br />

B: mandible and frons; C: pleotelson<br />

and uropod; D: pylopod; E: frons <strong>of</strong> praniza;<br />

F: first antenna, male.<br />

Synonyms: None.<br />

Diagnosis: Gnathia with a produced and entire<br />

frons, eyes lack<strong>in</strong>g. Mandible with a pronounced<br />

redan and three teeth. Pylopod with three articles,<br />

last article m<strong>in</strong>ute. Flagellum <strong>of</strong> first antenna with<br />

five articles, second also with five articles. Last two<br />

large peraeonal somites with a dorsal medial sulcus.<br />

Peraeopods not markedly sp<strong>in</strong>ous. Pleopoda lack<strong>in</strong>g<br />

setae.<br />

Measurements: Holotype male length 2.8 mm..<br />

width pleotelson 0.2 mm.<br />

Type locality: South <strong>Atlantic</strong>, L.G.O. Biotrawl No.<br />

55, type plus two male paratypes and one praniza.<br />

cat. no. 1-41.<br />

Distribution: Known only from <strong>the</strong> type locality.<br />

Aff<strong>in</strong>ities: This species is closely allied to Gnathia<br />

caeca Richardson, from which it differs <strong>in</strong> hav<strong>in</strong>g teeth<br />

on <strong>the</strong> mandible; o<strong>the</strong>rwise <strong>the</strong> two are very similar.<br />

Gnathia caeca Richardson<br />

Figure 3 A-B<br />

Synonyms: Gnathia caeca Richardson, 1911, pp.<br />

519-520; Stephensen, 1915, p. 7; Monod, 1926a,<br />

pp. 406-408.<br />

Diagnosis: Pylopods with three articles. Head as<br />

wide as thorax. Eyes lack<strong>in</strong>g. Frons produced but<br />

not as far as mandible apex. Supraocular lobes<br />

strongly tuberculate. Peraeopods without many stout<br />

sp<strong>in</strong>es. Fourth peraeonal somite with a pronounced<br />

dorso-medial sulcus. Marg<strong>in</strong>s <strong>of</strong> uropods and telson<br />

entire, without teeth.<br />

Measurements: Length 5 mm. (Monod, op. cit.,<br />

p. 407).<br />

Type locality: North <strong>Atlantic</strong>, Talisman Station 76,<br />

latitude 25° 1' N., longitude 19° 15' W., 2638 meters<br />

(Monod, op. cit., p. 408).<br />

Distribution: Known only from type locality.<br />

Aff<strong>in</strong>ities: <strong>The</strong> species appears to be most nearly<br />

related to G. bicolor Hansen <strong>in</strong> key characteristics at<br />

least [vide Monod, op. cit., p. 332).<br />

Gnathia stygia (G. O. Sars)<br />

Figure 3 C-E<br />

Synonyms: Anceus stygius G. O. Sars, 1877, p.<br />

348; — 1885, pp. 85-92; — 1886, pp. 27, 85. Gnathia<br />

stygius (G. O. Sars), Stebb<strong>in</strong>g, 1893, p. 338; —Ohl<strong>in</strong>,<br />

1901, p. 22, Fig. 3. Caecognathia stygia (G. O. Sars),<br />

Dollfus, 1901, p. 244. Caecognathia sarsi A. Dollfus,<br />

1901, pp. 244-245, Fig. 3. Bathygnathia stygia (err.<br />

typ.), Stephensen, 1915, p. 6. Caecognathia stygia<br />

(G. O. Sars), Stephensen, 1915, p. 7. Gnathia Stygia<br />

(G. O. Sars), Hansen. 1916, pp. 230-232; Monod,<br />

1926a, pp. 398-405.<br />

Diagnosis: Pylopods with three articles. Head<br />

narrower than thorax. Eyes lack<strong>in</strong>g. Frons produced

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