118 ROBERT J. MENZIES Figure 20. A-D: Haplomesus angustus Hansen. A: dorsal view immature male; B: ventral view male pleon; C: dorsal view male pleon; D: dorsal view head <strong>of</strong> male. E: Haplomesus modestus Hansen, dorsal view mutilated female. F-H: Haplomesus quadrisp<strong>in</strong>osus (G. O. Sars). F: ventral view male pleon from Ingolf Sta.. 36; G: dorsal view male cephalon from Ingolf Sta. 36; H: ventral view median lamella <strong>of</strong> male abdom<strong>in</strong>al operculum from Ingolf Sta. 36. I-J: Haplomesus tenuisp<strong>in</strong>is Hansen. I: dorsal view fragment <strong>of</strong> mutilated female from Ingolf Sta. 22; J: dorsal view <strong>of</strong> cephalon <strong>of</strong> female from Ingolf Sta. 24. K-M: Haplomesus tropicalis, n. sp. K: apex <strong>of</strong> male pleopod; L: dorsal view <strong>of</strong> male pleon and uropods; M: dorsal view male holotype. N-P: Haplomesus <strong>in</strong>signis Hansen. N: ventral view male pleon; O: dorsal view male cephalon; P: dorsal view male.
Haplomesus anguslus Hansen Figure 20 A-D Synonyms: Haplomesus anguslus. Hansen, 1916, pp. 61-62. PI, V. Diagnosis: Haplomesus with lateral sp<strong>in</strong>es on <strong>the</strong> first peraeonal somite only. Uropoda longer than wide. Pleon with pronounced postero-lateral angles dorsum without car<strong>in</strong>ae. Measurements: Juvenile male 4.8 mm. long. Full-grown males 9-12 mm. Length presumed (Hansen, op. cit.). Type locality: North <strong>Atlantic</strong>, south <strong>of</strong> Denmark Strait, Ingolf Station 18, latitude 61° 44' N., longitude 30° 29' W.; 1135 fathoms (2137 meters), temperature 3.0° C. (Hansen, op. cit.). Distribution: Also found by Ingolf Station 125, latitude 68° 08' N., longitude 16° 02'W., 729 fathoms (1373 meters), temperature —0.8° C. This species was not represented <strong>in</strong> Vema collections, which appear to lie too far south <strong>of</strong> <strong>the</strong> distributional area <strong>of</strong> <strong>the</strong> species. Gorbunov (1946, p. 123) reports this species <strong>in</strong> <strong>the</strong> Arctic from 698 meters. Aff<strong>in</strong>ities: This species superficially resembles S. gorbunovi Gurjanova from <strong>the</strong> Arctic; however, <strong>the</strong> postero-medial marg<strong>in</strong> <strong>of</strong> <strong>the</strong> pleon is trilobed ra<strong>the</strong>r than a s<strong>in</strong>gle lobe as <strong>in</strong> <strong>the</strong> latter. Haplomesus modestus Hansen Figure 20 E Synonyms: Haplomesus modestus. Hansen, 1916, pp. 65-66, PL V. Diagnosis: Haplomesus with lateral sp<strong>in</strong>es on somites 2, 3, and 4 and probably also on 1. Pleon without project<strong>in</strong>g postero-lateral angles, dorsum smooth and lack<strong>in</strong>g sp<strong>in</strong>es or car<strong>in</strong>ae. Uropoda not much longer than wide and not extend<strong>in</strong>g beyond posterior marg<strong>in</strong>. Measurements: 1.8 mm., fragment (Hansen, op. cit.). Type locality: North <strong>Atlantic</strong>, Davis Strait, Ingolf Station 24, latitude 63° 06' N., longitude 56° 00' W., 1199 fathoms (2258 meters), temperature 2.4° C., one mutilated female specimen (Hansen, op. cit.). Distribution: Known only from type locality. Aff<strong>in</strong>ities: <strong>The</strong> presence <strong>of</strong> lateral sp<strong>in</strong>es on peraeonal somites 2-4 <strong>in</strong>clusive dist<strong>in</strong>guishes this species from <strong>the</strong> o<strong>the</strong>rs that are known. Haplomesus quadrisp<strong>in</strong>osus (G. O. Sars) Figure 20 F-H Synonyms: Ischnosoma quadrisp<strong>in</strong>osum G. O. Sars, 1879, p. 435; — 1885, p. 126, PL II. Haplomesus quadrisp<strong>in</strong>osus (G. O. Sars), Richardson, 1908a, p. 81; — Hansen, 1916, pp. 59-61, PL II. ABYSSAL ATLANTIC ISOPODS 119 Diagnosis: Haplomesus with lateral sp<strong>in</strong>es on <strong>the</strong> first and third peraeonal somites only. Uropoda short, no longer than wide. Peraeon and pleon markedly tuberculate. Pleon without postero-lateral angles, dorsum with a s<strong>in</strong>gle medial swell<strong>in</strong>g. Apex <strong>of</strong> male first pleopods without lateral projections and with three term<strong>in</strong>al setae. Measurements: Three males, largest 5 mm. (after Hansen, op. cit.). Type locality: West <strong>of</strong> Norway, latitude 67° 56' N., longitude 4° 11' E., 1423 meters, temperature — 1.4°C (Hansen, op. cit.). Distribution: North <strong>Atlantic</strong>. <strong>The</strong> Ingolf collected it at n<strong>in</strong>e stations: Davis Strait: Station 24, latitude 63° 06' N., longitude 56° 00' W., 2258 meters, temperature 2.4° C., two specimens; Station 36, latitude 61° 50' N., longitude 56° 21' W., 2702 meters, temperature 1.5° C, two specimens. South <strong>of</strong> Davis Strait: Station 38, latitude 59° 12' N., longitude 51° 05' W., 3521 meters, temperature 1.3° C, one specimen. North <strong>of</strong> <strong>the</strong> Faeroes: Station 139, latitude 63° 36' N., longitude 7° 30'W., 1322 meters, temperature —0.6° C., two specimens. East <strong>of</strong> Iceland: Station 102, latitude 66° 23' N., longitude 10° 26' W., 1412 meters, temperature —0.9° C., five specimens. Nor<strong>the</strong>ast <strong>of</strong> Iceland: Station 120, latitude 67° 29' N., longitude 11° 32' W., 1666 meters, temperature — 1.0° C, one specimen; Station 119, latitude 67° 53' N., longitude 10° 19' W., 1902 meters, temperature —1.0° C., one specimen. South <strong>of</strong> Jan Mayen: Station 113, latitude 69° 31' N., longitude 7° 06' W., 2465 meters, temperature —1.0° C., five specimens; Station 116, latitude 70° 05' N., longitude 8° 26' W., 699 meters, temperature —0.4° C, one specimen. <strong>The</strong> species was not represented <strong>in</strong> <strong>the</strong> Lamont collections. Gorbunov (1946, p. 123) records it from 510 and 698 meters <strong>in</strong> <strong>the</strong> Arctic, and Birste<strong>in</strong> (1960, p. 15) cites it from <strong>the</strong> North Pacific at 4000-4150 meters. Aff<strong>in</strong>ities: This species resembles <strong>the</strong> new species ornatus considerably, but lacks paired car<strong>in</strong>ae on <strong>the</strong> dorsum <strong>of</strong> <strong>the</strong> pleon. Haplomesus tenuisp<strong>in</strong>is Hansen Figure 20 I-J Synonyms: Haplomesus tenuisp<strong>in</strong>is Hansen, 1916, pp. 64-65, PL V. Diagnosis: Haplomesus with lateral sp<strong>in</strong>es on peraeonal somites 1 and 4 only. Pleon with pronounced postero-lateral angles, dorsum with paired stout sp<strong>in</strong>es. Uropoda longer than wide and extend<strong>in</strong>g beyond <strong>the</strong> posterior border <strong>of</strong> <strong>the</strong> pleon.