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70<br />

BULLETIN OF THE UNIVERSITY OF NEBRASKA STATE MUSEUM<br />

crescent-shaped, sparsely setose or nearly<br />

glabrous area immediately behind margin<br />

<strong>of</strong> each mesocoxa. Metepimeron glabrous.<br />

Elytra: Each elytron with 2 broad, complete,<br />

transverse, orange bands. Humerus<br />

covered by field <strong>of</strong> short, stout setae that<br />

extend posteriorly to base <strong>of</strong> epipleural ridge<br />

(Figs. 85, 92). Epipleuron orange (Fig. 85).<br />

Legs: Anterior face <strong>of</strong> procoxa with sparse,<br />

minute setae on basal half. Posterior tibia<br />

straight. Posterior edge <strong>of</strong> metatrochanter<br />

with acute tooth projecting perpendicular<br />

from trochanter (Fig. 96).<br />

Distribution. Nicrophorus hybridus ranges<br />

from the southern part <strong>of</strong> western Canada<br />

southward through the north-central United<br />

<strong>State</strong>s to northern Arizona and New Mexico<br />

(Anderson and Peck 1985, Peck and Kaulbars<br />

1987). In <strong>Nebraska</strong>, this species is known<br />

from the northern portion <strong>of</strong> the state only<br />

but may also occur in the panhandle.<br />

Locality Records (Fig. 109). 3 <strong>Nebraska</strong><br />

specimens examined.<br />

CUMING CO. (2): West Point; SIOUX CO.<br />

(1): Warbonnet Canyon.<br />

Temporal Distribution. Rangewide: June<br />

to September (Peck and Kaulbars 1987).<br />

<strong>Nebraska</strong>: No data.<br />

Remarks. Nicrophorus hybridus is best identified<br />

by the combination <strong>of</strong> key characters,<br />

especially when trying to separate it from N.<br />

investigator. Nicrophorus hybridus is distinctive<br />

because <strong>of</strong> its broad elytral maculations,<br />

crescent-shaped region immediately behind<br />

each mesocoxa that is glabrous or sparsely<br />

setose, and stout humeral setae that extend<br />

to the base <strong>of</strong> the epipleural ridge. Both the<br />

males and females have small eyes. The posterior<br />

edge <strong>of</strong> the metatrochanter in the males<br />

has an acute tooth that projects perpendicular<br />

to the plane <strong>of</strong> the trochanter (Fig. 96). Contrast<br />

this with Fig. 97 for N. investigator.<br />

The larval stage was described by Anderson<br />

(1982), and a diagnosis was given by<br />

Anderson and Peck (1985).<br />

Little is known <strong>of</strong> the biology <strong>of</strong> this<br />

species. Peck and Kaulbars (1987) characterized<br />

their habitat as prairie, sage steppe, and<br />

montane meadow. Adults are probably diurnal<br />

and are reproductively active during the<br />

summer (Anderson and Peck 1985). Overwintering<br />

occurs in the prepupal stage.<br />

Nicrophorus investigator Zetterstedt<br />

(Figs. 17, 79, 86, 93, 97, 99, 101, 107, 110)<br />

Necrophorus investigator Zetterstedt 1824: 154.<br />

Necrophorus maritimus Guérin-Méneville<br />

1835: Pl. 17, Fig. 8.<br />

Necrophorus melsheimeri Kirby 1837: 97.<br />

Necrophorus particeps Fischer von Waldheim<br />

1844: 139.<br />

Necrophorus aleuticus Gistel 1848: 190.<br />

Necrophorus pollinctor Mannerheim 1853: 169.<br />

Necrophorus infodiens Mannerheim 1853: 170.<br />

Necrophorus confossor LeConte 1854: 20.<br />

Diagnosis. Length 13.2-18.0 mm. Head:<br />

Club <strong>of</strong> antenna with basal segment black,<br />

remaining segments orange. Male with small<br />

eyes, post-ocular bulge subequal to length<br />

<strong>of</strong> eye. Female with large eyes, post-ocular<br />

bulge less than half length <strong>of</strong> eye (Fig. 99).<br />

Thorax: Pronotum subquadrate, with lateral<br />

and basal margins wide; anterior, transverse<br />

depression deep (Fig. 79). Metasternum with<br />

dense yellow pubescence, including area just<br />

posterior <strong>of</strong> each mesocoxa. Metepimeron glabrous.<br />

Elytra: Pattern variable; each elytron<br />

with two broad, transverse, orange bands; or<br />

anterior band reduced to one, two, or three<br />

spots. Epipleuron orange (Fig. 86). Legs:<br />

Anterior face <strong>of</strong> procoxa with minute setae on<br />

basal half. Posterior tibia straight. Posterior<br />

edge <strong>of</strong> metatrochanter <strong>of</strong> males with acute<br />

tooth recurving dorsally (Fig. 97).<br />

Distribution. Nicrophorus investigator is<br />

widely distributed throughout Canada and<br />

Alaska and along the Rocky Mountains to<br />

New Mexico and Arizona. It is occasionally<br />

found in the northeastern United <strong>State</strong>s. It<br />

is also broadly distributed in Europe and<br />

Asia (Anderson and Peck 1985, Peck and

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