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Comparison of Stress and Learning Effects of Three Different ...

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Table 2.6 Behavioral elements <strong>of</strong> defensive threatening behavior developed from SCHENKEL (1947): (S), ZIMEN (1971):<br />

(Z), FEDDERSEN-PETERSEN u. OHL (1995): (F-O) <strong>and</strong> FEDDERSEN-PETERSEN (2004): (F), drawing on the example<br />

<strong>of</strong> ‘’Defensive-threat’’ from ZIMEN (1971)<br />

Facial Expression Signals<br />

Muzzle form long, pulled-back (Z) (F), poss. suddenly opening the mouth (Z) (F), at a sharp<br />

Corner <strong>of</strong> the mouth<br />

angle (F)<br />

Lips shape the lips being extremely drawn up or down (F),<br />

Nose bridge wrinkled (Z)<br />

Eyes/ Gaze direction<br />

Forehead<br />

baring the teeth (Z) (F)(S)<br />

Head posture lowered or lifted up against the opponent (Z)<br />

Ears position directed to forward (Z), the ear's root being pulled back or up (Z), lied back on<br />

the head (F),<br />

openings being plied together (F)<br />

Teeth bared all the way to the molars (F),<br />

Body Posture<br />

gingiva is visible (F)<br />

Signals<br />

Limbs/Joints crooked (Z) (F)<br />

Tail position tail drawn in (Z), With extreme body posture: tail pulled unto the belly (F)<br />

Tail activity stiff<br />

Hair piloerected on the back (Z) (F)<br />

Muscles<br />

Body posture hunch (Z), crouched (Z), slightly crouched (F)<br />

Other behavioral elements (Defensive-) snapping (S), uncompleted pouncing<br />

(S), pushing forward <strong>and</strong> biting (F), bite-clapping (S) (F),<br />

making oneself look small (F)<br />

Vocalizations silent (Z), hissing (Z), barking-growling (Z), growling-hissing (Z), growling (Z)<br />

(F), atonale barking (F-O)<br />

(F), squalling (F)<br />

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