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The Odonata of Sulawesi and adjacent islands ... - science . naturalis

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Libellago <strong>and</strong> Sclerocypha <strong>of</strong> <strong>Sulawesi</strong><br />

<strong>and</strong> a small marking in extreme posterior corner <strong>of</strong> mesepisternum; mesokatepisternum<br />

<strong>and</strong> mesepimeron black; metepisternum with extensive markings, a narrow<br />

zig-zag marking above metastigma, connected with subrectangular marking posterior<br />

to metastigma, <strong>and</strong> posterior to this marking an unconnected narrow crescentshaped<br />

marking next to interpleural suture; a large triangular marking in posterior<br />

corner <strong>of</strong> metepimeron.<br />

Abdomen: Narrow medio-dorsal pale line on S2-7, <strong>and</strong> a wider lateral stripe, covering<br />

S1-7, with spots on S8 <strong>and</strong> S9, interrupted on annulae.<br />

Measurements [mm]: Abdomen 15, Hw 20.<br />

Variation<br />

<strong>The</strong> most significant variation is in size.<br />

Measurements [mm]: male: Abdomen 16-18 mm, Hw 22-23.<br />

Biology<br />

Inhabits lowl<strong>and</strong> streams; males defend territories from boulders or emerging sticks<br />

in fast flowing water just above the water surface.<br />

Distribution<br />

Central <strong>Sulawesi</strong> around the northernmost part <strong>of</strong> the Teluk Bone between Palopo<br />

<strong>and</strong> Malili, south <strong>of</strong> the Kambuno <strong>and</strong> Balease mountains (Fig. 6); species <strong>of</strong> lowl<strong>and</strong><br />

rivers, not found at higher altitudes. Geographically isolated from L. c. celebensis by<br />

Tineba mountain range. L. c. anoa occurs in the mountains at higher elevations between<br />

400-600 m, <strong>and</strong> L. c. orientalis is found east <strong>of</strong> this area. Syntopic with L. rufescens<br />

near Palopo.<br />

Etymology<br />

Libellago celebensis orientalis ssp. nov.<br />

(Figs 2d, 3d, 4d, 5d, Plate IIc)<br />

Orientalis (Latin), eastern, because <strong>of</strong> its eastern distribution within <strong>Sulawesi</strong>. An<br />

adjective.<br />

Specimens examined<br />

All specimens in RMNH, unless stated otherwise. — Holotype P (JvT 5385): “1989<br />

RMNH Expedition to <strong>Sulawesi</strong>. Indonesia <strong>Sulawesi</strong> Tenggara: S <strong>of</strong> Sanggona: Mokowu<br />

river near Mokowu camp, foot <strong>of</strong> Watuwila Mts, 150 m asl, 3°48'S 121°39'E.<br />

Rivulet with ponded areas, clear water, bottom with boulders, some s<strong>and</strong>, largely<br />

shaded. Sample 89JvT031. 4 November 1989, Leg. J. van Tol”. With h<strong>and</strong>-written<br />

annotation in Dutch 'Gefotografeerd in verg. met L. ruf. van Centipede c.' [this speci -<br />

men was photographed in comparison with L. rufescens [recte: Watuwila vervoorti]<br />

from Centipede camp]. — Paratypes (147 specimens, per province in chronological<br />

order): <strong>Sulawesi</strong> Tengah. 1 P (JvT 6635), 15 km NNE <strong>of</strong> Luwuk: Sg. Biak, 150 m,<br />

International Journal <strong>of</strong> Odonatology 10 (2) 2007: 209-248, pls II, III<br />

227

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