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Revision of Lithostrotionella - USGS

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22 REVISION OF LITHOSTROTIONELLA FROM THE CARBONIFEROUS AND PERMIAN<br />

Diagnosis.- Cerioid Acrocyathus with corallite<br />

diameter 15 to 20 mm and 22 to 26 major septa that<br />

commonly reach the columella but seldom extend into<br />

the dissepimentarium. Minor septa absent or poorly<br />

developed. Columella seemingly simple to complex.<br />

Dissepimentarium composed <strong>of</strong> 1 to 3 rows <strong>of</strong> mostly<br />

large inflated dissepiments. Tabulae complete, conical,<br />

and spaced about 0.5 mm apart. Ratio <strong>of</strong> tabularium<br />

width to corallite diameter about 0.6. Mode <strong>of</strong> increase<br />

unknown.<br />

Description <strong>of</strong> holotype. -See Zhizhina (in Bul'vanker<br />

and others, 1960).<br />

Discussion. -At first glance, the transverse section <strong>of</strong><br />

this species suggests Stelechophyllum, but the tabulae<br />

are clearly <strong>of</strong> the Acrocyathus type, and the axial structures<br />

in some corallites appear to be complex.<br />

Occurrence. -Lower Carboniferous, middle or upper<br />

Visean. Zone C 1 vf, Donetz Basin, U.S.S.R.<br />

Acrocyathus cystosus (Zhizhina in Bul'vanker and others)<br />

Eolithostrotionella cystosa Zhizhina in Bul'vanker and others, 1960, p.<br />

250, pl. 61, figs. la, b.<br />

Diagnosis.- Cerioid Acrocyathus with corallite<br />

diameter 20 to 30 mm and 28 to 38 major septa that may<br />

or may not extend to the columella and seldom extend<br />

into the dissepimentarium. Minor septa poorly<br />

developed. Columella seemingly simple to complex.<br />

Dissepimentarium composed <strong>of</strong> as many as 5 rows <strong>of</strong><br />

dissepiments <strong>of</strong> varying sizes. Tabulae complete, conical,<br />

and spaced about 0.5 mm apart. Ratio <strong>of</strong>tabularium<br />

width to corallite diameter about 0.4 to 0.5. Mode <strong>of</strong> increase<br />

unknown.<br />

Description <strong>of</strong> holotype. - See Zhizhina (in Bul'vanker<br />

and others, 1960).<br />

Discussion. -This species is distinguished by its broad<br />

dissepimentarium.<br />

Occurrence.- Lower Carboniferous, lower to upper.<br />

Visean. Zones C 1<br />

vb to C 1<br />

vf, Donetz Basin, U.S.S.R.<br />

Acrocyathus lissitzini (Zhizhina in Bul'vanker and<br />

others)<br />

Eolithostrotionella lissitzini Zhizhina in Bul'vanker and others, 1960,<br />

p. 252, pl. 61, figs. 3a, b.<br />

Diagnosis.- Cerioid Acrocyathus with corallite<br />

diameter 15 to 20 mm and 18 to 25 major septa that approach<br />

the columella but seldom reach it and seldom extend<br />

into the dissepimentarium. Minor septa absent.<br />

Columella seemingly simple to complex. Dissepimentarium<br />

composed <strong>of</strong> 1 to 3 rows <strong>of</strong> dissepiments <strong>of</strong> varying<br />

sizes. Tabulae complete, conical, and spaced about 1<br />

mm apart. Ratio <strong>of</strong> tabularium width to corallite<br />

diameter about 0.4 to 0.5. Mode <strong>of</strong> increase unknown.<br />

Description <strong>of</strong>:holotype. -See Zhizhina (in Bul'vanker<br />

and others, 1960),.<br />

Occurrence. -Lower Carboniferous, middle or upper<br />

Visean. Zone C 1 vf, Donetz Basin, U.S.S.R.<br />

Acrocyathus hsujiulingi (Yoh)<br />

<strong>Lithostrotionella</strong> hsujiulingi Yoh, 1961, p. 8, 16, pl. 13, figs. la-c.<br />

Diagnosis.- Cerioid Acrocyathus with corallite<br />

diameter averaging 6 mm and about 16 major septa that<br />

approach the columella but seldom reach it and do not<br />

extend into the dissepimentarium. Minor septa well<br />

developed. Columella composed <strong>of</strong> a thickened axial<br />

plate joined to the counter septum, septal lamellae, and<br />

concentric traces <strong>of</strong> tabulae in transverse section.<br />

Dissepimentarium composed <strong>of</strong> a single row <strong>of</strong> dissepiments<br />

<strong>of</strong> varying sizes. Tabulae complete, conical,<br />

and spaced about 0.5 mm apart. Ratio <strong>of</strong> tabularium<br />

width to corallite diameter about 0.5. Mode <strong>of</strong> increase<br />

seemingly peripheral.<br />

Description <strong>of</strong> holotype.- See Yoh (1961).<br />

Discussion. -This species differs from the other<br />

Chinese species, A.? unicus, in having fewer major septa,<br />

fewer rows <strong>of</strong> dissepiments, and a somewhat more<br />

complex columella.<br />

Occurrence. -Lower Carboniferous(?), Kwangsi Province,<br />

China.<br />

Acrocyathus? unicus (Y abe and Hayasaka)<br />

Lithostrotion (<strong>Lithostrotionella</strong>) unicum Yabe and Hayasaka, 1915, p.<br />

133; 1920, p. 11, pl. 9, figs. 12a, b.<br />

<strong>Lithostrotionella</strong> unica Yabe and Hayasaka. Wu in Yii and others,<br />

1963, p. 86, pl. 24, figs. 7a, b.<br />

<strong>Lithostrotionella</strong> unicum Yabe and Hayasaka. Minato and Kato, 197 4,<br />

p. 72, pl. 15, fig. 1.<br />

Diagnosis. - Cerioid Acrocyathus? with corallite<br />

diameter averaging 7 mm and about 22 to 26 major septa<br />

that approach the columella but seldom reaeh it and<br />

seldom extend into the dissepimentarium. Minor septa<br />

well developed. Columella seemingly a simple axial plate<br />

augmented by a few axial tabellae and septal ]ameBae.<br />

Dissepimentarium composed <strong>of</strong> 1 to 3 rows <strong>of</strong> large inflated<br />

dissepiments. Tabulae complete, conical, and<br />

spaced 6 to 7 in 2 mm. Ratio <strong>of</strong> tabularium width to corallite<br />

diameter about 0.5. Mode <strong>of</strong> increase unknown.<br />

Description <strong>of</strong> holotype. -See Y abe and Hayasaka<br />

(1915, 1920) and Minato and Kato (1974).<br />

Discussion. -This species, the type <strong>of</strong> <strong>Lithostrotionella</strong>,<br />

is referred to Acrocyathus with query because<br />

<strong>of</strong> uncertainty concerning the morphology <strong>of</strong> the type<br />

specimen (see previous discussion on page 4).<br />

Occurrence.- Carboniferous(?), Yun-nan Province,<br />

China.<br />

Acrocyathus? shimeri (Crickmay)<br />

Lithostrotion pennsylvanicum Shimer, 1926, p. 27 [part].<br />

Lonsdaleia shimeri Crickmay, 1955, 1961, p. 13, pl. 1, figs. 9, 10.

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