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Revision of Lithostrotionella - USGS

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SYSTEMATIC PALE ONTOLOGY 31<br />

Discussion.-This species differs from P. wagneri by<br />

having more major septa, more tabulae, and strongly<br />

dilated corallite walls.<br />

Occurrence.- Middle Carboniferous, Bashkirian.<br />

PerapertU Formation, Palencia, Spain.<br />

Petalaxis radians (de Groot)<br />

<strong>Lithostrotionella</strong> (Hillia) radians de Groot, 1963, p. 89, pl. 20, figs.<br />

la-c.<br />

Diagnosis.-Petalaxis with corallite diameter 7 to 9<br />

mm and 15 to 18 major septa that closely approach the<br />

columella but seldom reach it and extend into the<br />

dissepimentarium, ordinarily being attached to the corallite<br />

wall. Minor septa ordinarily well developed. Columella<br />

a simple axial plate connected to the counter or<br />

cardinal septum. Dissepimentarium weakly lonsdaleoid,<br />

composed <strong>of</strong> 1 to 3 rows <strong>of</strong> inflated dissepiments <strong>of</strong> varying<br />

sizes. Tabulae incomplete, horizontal, mostly convex,<br />

and irregularly spaced 0.25 to 1 rrim apart. Ratio <strong>of</strong><br />

tabularium width to corallite diameter about 0.5 to 0.6.<br />

Mode <strong>of</strong> increase unknown.<br />

Description <strong>of</strong> type material.-See de Groot (1964<br />

[1963])..<br />

Discussion. -This species differs from P. wagneri by<br />

having larger corallites, more rows <strong>of</strong> dissepiments, and<br />

less regular tabulae.<br />

Occurrence.- Middle Carboniferous, Bashkirian.<br />

Perapertu Formation, Palencia, Spain.<br />

Petalaxis intermedius (de Groot)<br />

<strong>Lithostrotionella</strong> (Hillia) intermedia de Groot, 1963, p. 90, pl. 20, figs.<br />

2a, b, 3.<br />

Diagnosis. -Petalaxis with corallite diameter 5 to 7<br />

mm and 16 to 19 major septa that closely approach the<br />

columella but seldom reach it and extend into the<br />

dissepimentarium, commonly being attached to the corallite<br />

wall. Minor septa well developed. Columella a simple<br />

axial plate connected to counter or cardinal septum.<br />

Dissepimentarium weakly lonsdaleoid, CQmposed <strong>of</strong> 1 to<br />

4 rows <strong>of</strong> mostly small inflated dissepiments. Tabulae<br />

mostly incomplete, flat, concave or convex, horizontal,<br />

and irregularly spaced 0.25 to 0. 75 mm apart. Ratio <strong>of</strong><br />

tabularium width to corallite diameter about 0. 7 to 0.8.<br />

Increase peripheral.<br />

Description <strong>of</strong> type material.-See de Groot (1964<br />

[1963]).<br />

Discussion. -This species differs from P. wagneri by<br />

having larger corallites, more major septa, and more<br />

tabulae.<br />

Occurrence.- Middle Carboniferous,_ Bashkirian.<br />

Perapertu Formation, Palencia, Spain.<br />

Petalaxis santaemariae (de Groot)<br />

<strong>Lithostrotionella</strong> (Hillia) santaemariae de Groot, 1963, p. 91, pl. 21,<br />

figs. la-e.<br />

Diagnosis.-Petalaxis with corallite diameter 3.5 to 8<br />

mm and 16 to 19 major septa that closely approach the<br />

columella but seldom reach it and extend into the<br />

dissepimentarium, ordinarily attached to the corallite<br />

wall. Minor septa poorly developed. Corallite walls<br />

strongly dilated. Columella a simple thickened axial<br />

plate connected to counter or cardinal septum.<br />

Dissepimentarium weakly lonsdaleoid, composed <strong>of</strong> a<br />

single row <strong>of</strong> small inflated dissepiments. Tabulae mostly<br />

complete, horizontal, mostly concave, and irregularly<br />

spaced 0.25 to 1 mm apart. A peripheral zone <strong>of</strong><br />

clinotabellae may be present. Ratio <strong>of</strong> tabularium width<br />

to corallite diameter about 0.8. Increase peripheral.<br />

Description <strong>of</strong> holotype.- See de Groot (1964 [1963]).<br />

Discu,ssion.- This species differs from P. perapertuensis<br />

by having larger corallites, poorly developed minor<br />

septa, and fewer tabulae.<br />

OccU'rrence.- Middle Carboniferous, Bashkirian. Santa<br />

Maria Limestone, Palencia, Spain.<br />

.Petalaxis cantabricus (de Groot)<br />

<strong>Lithostrotionella</strong> (Hillia) cantabrica de Groot, 1963, p. 92, pl. 22, figs.<br />

1-4.<br />

Diagnosis. -Petalaxis with corallite diameter 4 to 8<br />

mm and 19 to 28 major septa that closely approach the<br />

columella but seldom reach it and extend into the<br />

dissepimentarium, ordinarily being attached to the corallite<br />

wall. Minor septa well developed. Cora!lite wall<br />

strongly dilated and denticulate. Columella a simple<br />

commonly thickened axial plate connected to counter or<br />

cardinal septum. Dissepimentarium weakly lonsdaleoid,<br />

composed <strong>of</strong> 1 to 3 rows <strong>of</strong> small inflated dissepiments.<br />

Tabulae mostly complete, horizontal, concave, and<br />

spaced 0.2!5 to 1 mm apart. A peripheral zone <strong>of</strong><br />

clinotabellae may be present. Ratio <strong>of</strong> tabularium width<br />

to corallite diameter about 0.6 to 0.7. Increase<br />

peripheral.<br />

Description <strong>of</strong> type material.- See de Groot (1964<br />

[1963]).<br />

Discussion. -This species is distinguished from all<br />

others in the P. wagneri species group by having more<br />

major septa.<br />

Occurrence.- Middle Carboniferous, Bashkirian to<br />

Moscovian(!'). Santa Maria Limestone, Perapertu Formation,<br />

and Celada Limestone, Palencia, Spain.<br />

Petalaxis orboensis (de Groot)<br />

<strong>Lithostrotionella</strong> orboensis de Groot, 1963, p. 85, pl. 17, figs. 2a-d.<br />

Diagnosis. -Petalaxis with corallite diameter 4.5 to 8<br />

'mm and 16 to 21 major septa that closely approach the<br />

columella but seldom reach it and extend into the<br />

dissepimentarium, commonly being attached to the corallite<br />

wall. Minor septa well developed. Columella a simple<br />

axial plate connected to counter or cardinal septum;

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