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<strong>Proceedings</strong> <strong>Anniversary</strong> <strong>Event</strong> <strong>–</strong><br />

<strong>10</strong> th <strong>International</strong> <strong>Alpine</strong> <strong>Workshop</strong><br />

Tagliamento 2012<br />

In Commemoration of Prof. Dr. Norbert Grosser's Retirement<br />

Compiled by<br />

Norbert Müller & Norbert Grosser & Martin Kümmerling<br />

University of Applied Sciences Erfurt, Landscape Architecture (Ed.)<br />

Erfurt 2012<br />

1


<strong>Proceedings</strong> <strong>Anniversary</strong> <strong>Event</strong> <strong>–</strong><br />

<strong>10</strong> th <strong>International</strong> <strong>Alpine</strong> <strong>Workshop</strong><br />

Tagliamento 2012<br />

In Commemoration of Prof. Dr. Norbert Grosser's Retirement<br />

Dear Norbert Grosser,<br />

we want to dedicate this special <strong>Anniversary</strong> <strong>Proceedings</strong> in commemoration of your<br />

retirement from University.<br />

We wish to express our deep acknowledgement for guiding us to animal diversity of the<br />

wonderful Tagliamento River and hope that we will have the chance to participate from<br />

your rich experience on future workshops, too.<br />

Your colleagues, students and former students<br />

2


CONTENT<br />

1 GENERAL INFORMATION ON THE INTERNATIONAL ALPINE WORKSHOP <strong>–</strong> TAGLIAMENTO 2012 ..4<br />

1.1 ANNOUNCEMENT & PROGRAMME............................................................................................ 4<br />

1.2 ORGANISERS & PARTICIPANTS ............................................................................................... 6<br />

2 GENERAL INFORMATION ABOUT THE ALPS & THE ALPINE FORELAND (NORBERT MÜLLER) ......7<br />

2.1 GENESIS & ECOLOGY............................................................................................................. 7<br />

2.2 ALTITUDINAL BELTS, PLANT COMMUNITIES & LIVE FORMS IN THE ALPS ................................... 12<br />

3 ECOLOGY & VEGETATION OF ALPINE RIVERS (NORBERT MÜLLER) .......................................14<br />

3.1 FLOODPLAIN MORPHOLOGY.................................................................................................. 14<br />

3.2 CHARACTERISTICS OF PLANTS AND PLANT COMMUNITIES ...................................................... 15<br />

3.3 ALTERATION OF THE FLOODPLAIN VEGETATION UNDER HUMAN IMPACT .................................. 17<br />

3.4 SITUATION OF KEY SPECIES IN ALPINE FLOODPLAINS ............................................................. 19<br />

4 INVESTIGATION AREAS & FFH SITES ALONG THE TAGLIAMENTO RIVER.................................20<br />

4.1 INVESTIGATION AREA 1 <strong>–</strong> PASSO DELLA MAURIA ................................................................... 21<br />

4.2 FFH SITE IT3230089 <strong>–</strong> DOLOMITI DEL CADORE E DEL COMELICO.......................................... 22<br />

4.3 INVESTIGATION AREA 2 <strong>–</strong> FORNI DI SOTTO ............................................................................ 23<br />

4.4 FFH SITE IT33<strong>10</strong>001 <strong>–</strong> DOLOMITI FRIULANE ........................................................................ 24<br />

4.5 INVESTIGATION AREA 3 <strong>–</strong> AMARO / FELLA ............................................................................. 26<br />

4.6 FFH SITE IT3320015 <strong>–</strong> VALLE DEL MEDIO TAGLIAMENTO ..................................................... 27<br />

4.7 INVESTIGATION AREA 4 <strong>–</strong> CORNINO & MONTE DI RAGOGNA ................................................... 28<br />

4.8 FFH SITE IT33<strong>10</strong>007 <strong>–</strong> GRETO DEL TAGLIAMENTO ............................................................... 29<br />

4.9 INVESTIGATION AREA 5 <strong>–</strong> SPILIMBERGO ................................................................................ 30<br />

4.<strong>10</strong> INVESTIGATION AREA 6 <strong>–</strong> CASARSA ...................................................................................... 31<br />

4.11 INVESTIGATION AREAS 7 & 8 <strong>–</strong> BOLZANO & LATISANA............................................................ 31<br />

4.12 INVESTIGATION AREA 9 <strong>–</strong> BIBIONE / ESTUARY ....................................................................... 32<br />

4.13 FFH SITE IT3250033 <strong>–</strong> LAGUNA DI CAORLE / FOCE DEL TAGLIAMENTO ................................. 32<br />

4.14 INVESTIGATION AREA <strong>10</strong> <strong>–</strong> FLAMBRO .................................................................................... 33<br />

4.15 FFH SITE IT3320026 <strong>–</strong> RISORGIVE DELLO STELLA ............................................................... 34<br />

4.16 INVESTIGATION AREA 11 <strong>–</strong> VIVARO ....................................................................................... 35<br />

4.17 FFH SITE IT33<strong>10</strong>009 <strong>–</strong> MAGREDI DEL CELINA ...................................................................... 35<br />

5 METHODS OF RESEARCH ..................................................................................................37<br />

5.1 ASSESSMENT OF VASCULAR PLANTS, KEY SPECIES & NEOPHYTES (NORBERT MÜLLER).......... 37<br />

5.2 RESEARCH PROGRAMME ANIMAL ECOLOGY (NORBERT GROSSER) ........................................ 39<br />

6 POSTER SERIES "TAGLIAMENTO <strong>–</strong> KING OF ALPINE RIVERS" ..............................................40<br />

7 FLORA OF THE TAGLIAMENTO (NORBERT MÜLLER) ............................................................56<br />

8 LITERATURE ....................................................................................................................67<br />

8.1 GENERAL LITERATURE: ........................................................................................................ 67<br />

8.2 WORKING LITERATURE AND MAPS......................................................................................... 69<br />

9 LEGEND TO PICTURES ON FRONT & BACK COVER................................................................70<br />

3


1 GENERAL INFORMATION ON THE INTERNATIONAL ALPINE WORKSHOP<br />

<strong>–</strong> TAGLIAMENTO 2012<br />

1.1 ANNOUNCEMENT & PROGRAMME<br />

4


Appendix 1<br />

Program "<strong>10</strong> th Year <strong>Anniversary</strong> <strong>Event</strong> <strong>–</strong> 2 nd River Conference Tagliamento <strong>–</strong><br />

Research and education at alpine rivers"<br />

Location Hotel Ai Glicini, Cornino (Friaul), Italy<br />

Time 14.00 <strong>–</strong> 19.00<br />

14.00 Welcome<br />

14.<strong>10</strong> History of <strong>Alpine</strong> <strong>Workshop</strong>s at the Tagliamento River <strong>–</strong> Norbert Müller <strong>–</strong> Dep.<br />

Landscape Management & Restoration Ecology, University of Applied Sciences<br />

Erfurt<br />

15.00 An introduction to the work of WWF at the Tagliamento and in the Alpes <strong>–</strong> Mauro<br />

Belardi <strong>–</strong> WWF Italy & WWF European <strong>Alpine</strong> Programme<br />

15.30 Discussion and coffee break<br />

16.30 Review on the Tagliamento <strong>Workshop</strong> 2005 <strong>–</strong> Martin Kümmerling <strong>–</strong> Dep.<br />

Landscape Management & Restoration Ecology, University of Applied Sciences<br />

Erfurt<br />

17.00 Tagliamento River area as a habitat complex of special faunal interest and value <strong>–</strong><br />

Norbert Grosser <strong>–</strong> Dep. Zoology & Animal Ecology, University of Applied Sciences<br />

Erfurt<br />

17.30 Ecology and vegetation of braided rivers in the European Alps <strong>–</strong> Norbert Müller <strong>–</strong><br />

Dep. Landscape Management & Restoration Ecology, University of Applied<br />

Sciences Erfurt<br />

18.00 Ecological and cultural river landscape management on the Tango Peninsula,<br />

Kyoto, Japan <strong>–</strong> Katsue Fukamachi & Iwao Miyoshi <strong>–</strong> Kyoto University, Japan<br />

18.30 Discussion and final statement<br />

19.00 Press release and dinner at Ai Glincini<br />

Appendix 2 & 3<br />

Research program Fauna Flora Habitats<br />

Please see chapter 5 <strong>–</strong> Methods of Research.<br />

5


1.2 ORGANISERS & PARTICIPANTS<br />

Name duties phone e-mail<br />

Prof. Dr. Grosser,<br />

Norbert<br />

Prof. Dr. Müller,<br />

Norbert<br />

Chairs University of Applied Sciences Erfurt<br />

- responsible animal ecology 1624139111 grosser@fh-erfurt.de<br />

- overall coordinator,<br />

- responsible vegetation ecology,<br />

- hotel reservation for Erfurt participants<br />

6<br />

1795123761 n.mueller@fh-erfurt.de<br />

Scientific Assistants University of Applied Sciences Erfurt<br />

Gabler, Sebastian<br />

- animal ecology<br />

BA Student<br />

König, Sebastian<br />

BA Student<br />

Kümmerling, Martin<br />

PhD student<br />

Amschler, Julia<br />

- Contact person external participants<br />

- Bus reservation 1<br />

- vegetation ecology<br />

- vegetation ecology<br />

- proceedings/excursion guide<br />

- hardware for conference<br />

15208950525<br />

1797503058<br />

Students University of Applied Sciences Erfurt (BA 4. Semester)<br />

Bachmann, Helena - herbarium 1706640008<br />

Bettin, Felix<br />

- Contact person Erfurt students<br />

- Bus reservation 2<br />

- driver 1 Bus 2<br />

sebastian.koenig@fherfurt.de <br />

martin.kuemmerling@fherfurt.de <br />

Julia.amschler@fherfurt.dehelena.bachmann@fherfurt.de<br />

15118180344 felix.Bettin@fh-erfurt.de<br />

Kiesel, Torben - students' present 15151632244<br />

Torben.kiesel@fherfurt.de<br />

Krautz, Stephanie<br />

- driver 2 Bus 2<br />

- treasurer<br />

17665707536<br />

Stephanie.krautz@fherfurt.de<br />

Kühn, Robert 16090229121 Robert.kuehn@fherfurt.de<br />

Kupfer, Katharina - literature 17696575807<br />

Katharina.kupfer@fherfurt.de<br />

Mainka, Matthias - driver 1 Bus 1 1757987040<br />

Matthias.mainka@fherfurt.de<br />

Rembold, Matthias - driver 2 Bus 1 17670804482 Matthias.rembold@remat.<br />

de<br />

Schollän, Christian 15117720857 Christian.schollaen@stud.<br />

fh-erfurt.de<br />

Schuberth, Viktoria<br />

Wieja, Stefanie<br />

- herbarium<br />

- treasurer<br />

1773719556<br />

Students Technical University Berlin (4. Semester)<br />

Stefanie.wieja@fherfurt.de<br />

Abraham, Stefanie 017632891154 st_abraham@arcor.de<br />

Ruockute, Ruta 015782386700 rutaruo@gmail.com<br />

Wolf, Kathrin 01636145630 kathrinwolf89@yahoo.de<br />

Wunder, Tobias 01773779767<br />

Belardi, Mauro<br />

WWF Italy<br />

Vespermann, André<br />

FH Erfurt (Alumni)<br />

Researchers & Guests<br />

t-wunder@mailbox.tuberlin.de<br />

m.belardi@wwf.it<br />

01797534621 vesper-man@gmx.de


2 General information about the Alps & the <strong>Alpine</strong> foreland (Norbert<br />

Müller)<br />

2.1 GENESIS & ECOLOGY<br />

About 200 Million years ago the supercontinent Pangaea broke (fig. 2) in two blocks:<br />

Laurasia in the north and Gondwana in the south. The propulsions of this earth's crust<br />

movement are slowly circulating convection streams in lower, plastically layers of the<br />

earth's mantle. The heavier ocean ground is pulled in the depth, whilst the lighter<br />

continental massif float with an average speed of 2 cm per year and were upheaved to<br />

bedrocks along the seam line.<br />

Differently the seam lines between two broken continental massifs: now the lava current<br />

raise from the depth nearly up to the earth or ocean crust. Long ridges traced back by<br />

volcanism activities are the result (Mid-Atlantic ridge). Caused by the collision between the<br />

African and the Eurasian plate, the ground of the Tethys <strong>–</strong> the old Mediterranean basin <strong>–</strong><br />

has been pressed together and resulted in overlays, rifts, uplifts and convolution (fig. 3).<br />

7


The origin of the Alps (and the entire mountain range from the High Atlas over the<br />

Himalaya to New Zealand) started about 40 Million years ago. At the same time ocean<br />

sediments were also raised or transported as whole layer packages and partially pressed<br />

in folds. In several impulses a low mountain range relief was formed firstly; only in a<br />

younger geologic era, during the last 2 Million years, the Alps have become the high<br />

mountains with its sharp erosion patterns.<br />

The formation of mountains as well as the denudation are current processes. The present<br />

rock structures of the eastern part of the Alps shows a clear longitudinal symmetry: The<br />

central silicate mountains originated from the ancient magmatic earth crust were flanked<br />

north- and southwards by limestone ranges. Those result from sea sediments or reefs of<br />

calcareous algae and corals which were emerged in the tropical shallow sea of the Trias<br />

period in the Tethys and grew up slowly to mighty rock ‘castles’ (especially the Dolomites).<br />

Sometimes younger volcanic activities have disturbed the calm development of this ocean<br />

area (Bozner quartz- porphyry plate). The western Alps are shifted and folded much more<br />

complex and limestone sediments, shale and siliceous primary rocks change irregularly on<br />

a close area (fig. 2 - 4 from Reisigl & Keller 1987).<br />

Fig. 2: Starting continental drift<br />

end of Permian (225 Mill. years<br />

ago)<br />

Fig. 5: The Wettersteingebirge, like the western<br />

parts of the Dolomites in South Tirol, belongs to a<br />

big reef area of the Mesozoic era.<br />

The rocks were built in Trias by the skeletons of<br />

the corals or they were deposited in shallow<br />

lagoons.<br />

The figure shows top view and profile of a<br />

calcareous alga (Typus Triploporella, 2x). Outside<br />

the living alga cells and inside the calcareous<br />

skeleton.<br />

Fig: 3: Drift phase end of<br />

Cretaceous (65 Mill. years ago)<br />

starting mountain building<br />

process<br />

8<br />

Fig. 4: Profile through collision<br />

zone


Fig. 6: Fold of mountain formation. Folds at the<br />

mountain surface are partly removed by erosion.<br />

The former stratification and the age of the single<br />

layer is mostly reconstructed by "index fossils".<br />

Fig. 7: The siliceous mountains, like the Ötztaler<br />

Alps, existing of primary rocks. In the Palaeozoic,<br />

but also in geologically younger times (Tertiary)<br />

liquid magma emerged from earth’s interior and<br />

forms as granite or basalt the base of the<br />

continental massifs, whicher later were covered<br />

with a slight layer of marine deposits.<br />

Profile through a young volcano which had broken<br />

through the ocean sediments.<br />

(Fig. 5-7 from Reisigl & Keller 1987)<br />

Fig 8: Tectonic overview of Europe (from Westermann 1991)<br />

9


The flora of the Northern and Central Alps is relatively young, like in other nearly<br />

completely glaciated high mountains. In contrast to the massif of Southern Alps, which<br />

were ice-free during cold stages, these parts only accommodate few endemic species.<br />

Nevertheless, the essential <strong>Alpine</strong> flora (above the forest line) is relatively independent<br />

compared with those of the <strong>Alpine</strong> Foreland.<br />

Beside modified and separate forms of lowland phanerogams many typical high mountain<br />

plants occur in the alpine level, which are originated in Asian and Southern European<br />

mountains as well as in the Arctic.<br />

Fig. 9: Climate and forest development in the<br />

postglacial period<br />

(from Reisigl & Keller 1987)<br />

Fig. <strong>10</strong>: Change of the flora character by climate<br />

change during the Tertiary:<br />

1. tropical elements, 2. Mediterranean<br />

elements, 3. temperate flora, 4. arctic &<br />

alpine elements<br />

(from Reisigl & Keller 1987)<br />

The northward exposed plateau between the Alps and the Danube River is called <strong>Alpine</strong><br />

Foreland. Its origin is strong related with the formation of the Alps.<br />

The more the Alps were lifted up and eroded, the more the foreland subsided and were<br />

filled with depositions brought by the mountain rivers. The entire tertiary debris of the Alps<br />

in its foreland area is summarized as molasses. The molasses is layered flat in the north,<br />

whilst in the south it was formed in some saddles and depressions and overlooks the<br />

landscape rib-like, e.g. near Murnau. During the ice age the glaciers expanded and gave<br />

the foreland its present diverse appearance.<br />

The ‘Inn Glacier’ covered during its peak level an area of approx. 1600 km² of the <strong>Alpine</strong><br />

foreland. In front of the alpine cross- valleys from where the glaciers originated, glacier<br />

tongue basins with finger-like branch basins are situated. In between undulating glacial<br />

drifts or tills and drumlin groups pervade the landscape. The tongue basins and also some<br />

<strong>10</strong>


anch basins contain lakes (Ammersee, Starnberger See, Kochelsee, Chiemsee). Lech,<br />

Isar, Inn and Salzach River have already lost their lakes. The borders of ancient glaciers<br />

are marked by the end moraine walls which surround the basins in a large arrow.<br />

Altitude (average 500 m NN) and distance to the Alpes determine the climate of the <strong>Alpine</strong><br />

forelands. Referring to the latitude the typical west weather conditions of Central Europe<br />

are strongly modified by the nearness of the high mountains in the <strong>Alpine</strong> foreland. During<br />

a southwest stream the Foehn wind conditions arise with dry and warm weather; during a<br />

northwest stream there are strong cloud accumulations as well as heavier and longer rain<br />

intensity. The cold snaps thereby hold in southern Bavaria twice as long as in alpinedistant<br />

areas (WESTERMANN 1962).<br />

Fig. 11: Europe during the last glacial age (from Westermann 1991)<br />

11


2.2 ALTITUDINAL BELTS, PLANT COMMUNITIES & LIVE FORMS IN THE ALPS<br />

Fig. 12: Altitudinal belts and plant communities on limestone <strong>–</strong> Northern & Southern Alps<br />

(from Reisigl & Keller 1987)<br />

12


Fig. 13: Altitudinal belts and plant communities on silicate <strong>–</strong> Central Alps<br />

(from Reisigl & Keller 1987)<br />

13


3 ECOLOGY & VEGETATION OF ALPINE RIVERS (NORBERT MÜLLER)<br />

3.1 FLOODPLAIN MORPHOLOGY<br />

River floodplains are habitats which originate from to fluvial processes and either<br />

influenced by flood occasions (present floodplain) or were flooded in former times (fossil<br />

floodplain).<br />

Rivers and their floodplains have varying responsibilities in the balance of nature: Run-off<br />

and retention of the precipitation water, evacuation of deposition material out of the<br />

mountains. For the protection of endangered species and biotope conservation river<br />

floodplains are of outstanding relevancy as central biotope linking axes and centres of<br />

biodiversity.<br />

River-morphologically there are three different river course types (Fig. 14, left to right):<br />

- stretched course (upper reaches, erosion predominates)<br />

- braided course (middle reaches, assembly-line balance between erosion and<br />

sedimentation)<br />

- bended course (low land, sedimentation predominates)<br />

Fig. 14: River morphology types<br />

14


3.2 CHARACTERISTICS OF PLANTS AND PLANT COMMUNITIES<br />

Floodplain specialists like German Tamarisc (Myricaria germanica) and Willow species<br />

(Salix div. spec.) are adapted to:<br />

a) stochastic disturbances (periodical or episodic floods)<br />

b) change of flooding and draughts<br />

- fast occupation of new habitats i.e. quick germination and propagation by wind and<br />

water<br />

- long root system as fixation and<br />

- high regeneration ability<br />

Adaptations of fauna: for instance pink-winged grasshopper (Bryodema tuberculata)<br />

Fig. 15: Phyto-sociological classification of the floodplain vegetation in Central Europe<br />

(from Pfadenhauer 1993)<br />

15


Head waters of alpine rivers <strong>–</strong> streched river course<br />

- Open woodless floodplain mostly without vegetation or with single pioneer vegetation<br />

from the alpine screes (K Thlaspietea rotundifolii)<br />

Upper reaches of alpine rivers <strong>–</strong> braided river course, inner alpine<br />

- Woodless floodplain with gravel bars; pioneer vegetation <strong>–</strong> K Thlaspietea rotundifolii <strong>–</strong><br />

beside floodplain species of the sub-alpine screes and alpine meadow communities<br />

- Softwood alluvial forests on the coarse gravel bars with single willows and willowtamarisk<br />

bush land (K Salicetea purpureae)<br />

- Hardwood alluvial forest consisting of alder-pine forests (close to groundwater) and<br />

winter heath-pine forests (K Erico-Pinetea) (fossil floodplain)<br />

Middle reaches of alpine rivers <strong>–</strong> braided river course, outer alpine<br />

- Open floodplain with sand and debris-rich islands; pioneer vegetation on gravel mixed<br />

with ruderal species (K Artemisietea)<br />

- Softwood alluvial forests (nearly above the mean water level) on coarse debris: willow<br />

bush land (A Salicetum eleagni)<br />

- On sandy sediments: willow-tamarisk bush land (K Salicetea purpureae),<br />

- River distant: alder forests (K Querco-Fagetea, A Alnetum incanae) and willow forests (K<br />

Salicetea purpurea, V Salicion albae)<br />

Silt-up vegetation of old river bends<br />

- Freshly developed with calcareous fen community e.g. K Scheuchzerio-Caricetea nigrae,<br />

A Equiseto-Typhetum minimae); in the further succession process substituted by reeds<br />

and bigger sedges (K Phragmitetea); calcareous-oligotroph waters with submerse<br />

Characeae<br />

- Hardwood alluvial forests (above the average high water level) sedimentation of fine<br />

material (sand, silt): oak-elm forests (K Querco-Fagetea, A Querco-Ulmetum); in the<br />

fossil floodplain on freshly developed gravel terraces sea buckthorn thickets and<br />

succession to winter heath-pine forests (K Erico-Pinetea)<br />

upper and middle reaches of low mountain range rivers (braided)<br />

- Woodless floodplain with gravel and sand bars <strong>–</strong> flood vegetation with winter cress<br />

community (K Artemisietea) and reeds on gravel (K Phragmitetea)<br />

16


- Softwood alluvial with willow bush land (K Salicetea purpureae, A Salicetum triandraeviminalis)<br />

and elm-ash forests (K Querco-Fagetea, A Pruno-Fraxinetum)<br />

Lower course of alpine rivers and low mountain range rivers (lowland floodplains) <strong>–</strong><br />

bended river course with slip-off slopes<br />

- Open floodplain with gravel and sand bars <strong>–</strong> flood vegetation winter cress community (K<br />

Artemisietea) and reeds on gravel (K Phragmitetea) and thermophyts (K Bidentetea) on<br />

sand and mud<br />

- Backwater with reeds and bigger sedges (K Phragmitetea)<br />

- Softwood alluvial forest with willows (K Salicetea pupureae)<br />

- Hardwood alluvial forest with elm-ash forests, oak-elm forests (all K Querco-Fagetea)<br />

3.3 ALTERATION OF THE FLOODPLAIN VEGETATION UNDER HUMAN IMPACT<br />

The influence of humans on flora and vegetation of big lowland floodplains, which means<br />

rivers with main catchments area in the low mountain ranges, goes back to a long time.<br />

Since the Neolithic Age the run-off characteristic and the bed load of the big lowland rivers<br />

were already changed by forest clearing in the catchments area. The intensive human<br />

impact on flora and vegetation of the lowland floodplains already started in the Roman's<br />

time by expansion the agriculture. The sedimentation of floodplain loam in the Weser<br />

valley, for instance, refers to the expansion of agriculture activities in big parts of the loesscovered<br />

hills in the catchments area. Before that gravel soils prevailed in the Weser<br />

floodplains and gave a completely different character.<br />

Bigger changes in the catchments area of the alpine rivers first happened since the Middle<br />

Ages. The stronger settlement development in the alpine valleys and the clearing of<br />

mountain forests caused a temporary raise of the bed load which led, for instance, to the<br />

creation of river terraces in the lower course of the Isar River (cf. Müller 1995).<br />

From the 18 th century onward mankind started to change many lowland floodplains and<br />

from the 19 th century the alpine floodplains were altered essentially.<br />

River regulation means forcing a branched river to run through a regulated channel by<br />

cross- and lengthwise constructions. Subsequent bed erosion was tried to countervail with<br />

the installation of bed ramps.<br />

Effects: strong decline of the open floodplains, softwood alluvial forests and backwaters,<br />

loss of the floodplain specialists like Typha minima (species that were adapted to natural<br />

17


disturbances), transformation of the hardwood alluvial forests into plantations and<br />

agricultural land.<br />

Construction of barrages is done since the middle of the 20 th century as a more energyefficient<br />

development (alpine rivers) or shipping extension (low mountain range rivers).<br />

Effects: Entire loss of the typical floodplain habitats by loss of the floodplain dynamics (loss<br />

of morpho-dynamic, altered hydrodynamics and discharge level by humans, loss of the<br />

ecological connectivity, strong consequences also on remaining water courses below<br />

water reservoirs).<br />

Fig.15: River dynamics and structure of floodplain vegetation (simplified depiction) at the braided course of a debris-rich<br />

northern alpine river (alpine foreland) before river regulation (from Müller 1995).<br />

Fig. 16: River dynamics and structure of floodplain vegetation (simplified depiction) at the lower course of a<br />

debris-rich northern alpine river (alpine foreland) after river regulation (from Müller 1995).<br />

18


3.4 SITUATION OF KEY SPECIES IN ALPINE FLOODPLAINS<br />

Fig. 17: Distribution of German Tamarisc (Myricaria germanica) along northern alpine rivers<br />

(from Müller 1995)<br />

19


4 INVESTIGATION AREAS & FFH SITES ALONG THE TAGLIAMENTO<br />

RIVER<br />

Fig. 18: Description of the drainage area, geology and watercourses with the investigation areas and<br />

FFH sites (from Lippert et al. 1995, changed Müller 2012)<br />

20


4.1 INVESTIGATION AREA 1 <strong>–</strong> PASSO DELLA MAURIA<br />

Fig. 19: Location of both investigation areas at Mauria (TK 25, Dolomiti di Friulane)<br />

Short description of the investigation area 1 Passo della Mauria (from Lippert et al. 1995):<br />

The Tagliamento rises near the Passo della Mauria, on a height of 1700 m above sea<br />

level. The investigation area lies around 500 m below the spring. In this section, the<br />

Tagliamento is a small mountain torrent with a strong slope. The brook bed consists of<br />

boulders, stones and slip rock. The varying water levels and the big amount of scree<br />

material cause that plants are either rare or just in initial stages.<br />

Appearing plant communities:<br />

- Thlaspietea<br />

- Salicetum elaeagni<br />

- Juncetum alpini<br />

- Pinus mugo-communities<br />

- Mixed woodland<br />

21


4.2 FFH SITE IT3230089 <strong>–</strong> DOLOMITI DEL CADORE E DEL COMELICO<br />

Fig. 20: Delimitation of the FFH-Area Dolomiti del Cadore e del Comelico (European Environmental Agency<br />

2009)<br />

Territory size: 70396.0000 ha Biogeographic region: alpine<br />

Altitude maximum: 3000 m Altitude minimum: 496 m<br />

Tab. 1: European Union Habitats and their percentage of the area (European Environmental Agency)<br />

Code EUH Description<br />

22<br />

Percentage<br />

of area<br />

3220 <strong>Alpine</strong> rivers and the herbaceous vegetation along their banks 1%<br />

3230 <strong>Alpine</strong> rivers and their lingeous vegetation with Myricaria germanica 1%<br />

3240 <strong>Alpine</strong> rivers and their lingeous vegetation with Salix eleagnos 1%<br />

4060 <strong>Alpine</strong> and Boreale heaths 3%<br />

4070* Bushes with Pinus mugo and Rhododendron hirsutum (Mugo-<br />

Rhododendretum hirsuti)<br />

8%<br />

4080 Sub-arctic Salix spp. scrub 2%<br />

6150 Siliceous alpine and boreal grasslands 4%<br />

6170 <strong>Alpine</strong> und subalpine calcareous grasslands 4%<br />

62<strong>10</strong> Semi-natural dry grasslands and scrubland facies in calcareous<br />

substrates (Festuco-Brometalia)<br />

1%<br />

6230*<br />

64<strong>10</strong><br />

6430<br />

Species-rich Nardus grasslands, on siliceous substrates in mountain<br />

areas (and submountain areas, in Continental Europe)<br />

Molinia meadows on calcareous, peaty or clayey-silt-laden soils<br />

(Molinion caeruleae)<br />

Hydrophilous tall herb fringe communities of plains and of the montane to<br />

alpine levels<br />

65<strong>10</strong> Lowland hay meadows (Alopecurus pratensis, Sanguisorba officinalis) 1%<br />

6520 Mountain hay meadows 1%<br />

1%<br />

1%<br />

2%


71<strong>10</strong>* Active raised bogs 1%<br />

7140 Transition mires and quaking bogs 1%<br />

7150 Depressions on peat substrates of the Rhynchosporion 1%<br />

7230 Alkaline fens 1%<br />

7240* <strong>Alpine</strong> pioneer formations of Caricion bicoloris-atrofuscae 1%<br />

81<strong>10</strong> Siliceous scree of the montane to snow levels (Androsacetalia alpinae<br />

and Galeopsietalia ladani)<br />

1%<br />

8120<br />

Calcareous and calcshist screes of the montane to alpine levels<br />

(Thlaspietea rotundifolii)<br />

82<strong>10</strong> Calcareous rocky slopes with chasmophytic vegetation 9%<br />

8220 Siliceous rocky slopes with casmophytic vegetation 1%<br />

8240* Limestone pavements 1%<br />

9130 Asperulo-Fagetum beech forests 3%<br />

9150 Medio-European limestone beech forests of the Cephalanthero-Fagion 1%<br />

9180* Tilio-Acerion forests of slopes, screes and ravines 1%<br />

91D0* Bog woodland 1%<br />

94<strong>10</strong> Acidophilous Picea forests of the montane to alpine levels (Vaccinio- 39%<br />

9420 <strong>Alpine</strong> Larix decidua and/or Pinus cembra forests 1%<br />

9530* (Sub-) Mediterranean pine forests with endemic black pines 4%<br />

* Priority habitat<br />

4.3 INVESTIGATION AREA 2 <strong>–</strong> FORNI DI SOTTO<br />

Short description of the investigation area 2 Forni di Sotto (from Lippert et al. 1995):<br />

In the mountains, the Tagliamento is characterized by a change between tight and wide<br />

river segments. The tight parts show a high transport of scree, while the wide segments<br />

are dominated by deposition and rearrangements. Because of this, the river creates an<br />

own riverbed. The investigation area is located underneath the Passo della Mauria on a<br />

height of 670 m above sea level. It is predestinated for the study of succession and zoning<br />

of floodplains.<br />

Pioneer vegetation:<br />

Chondrilletum chondrilloidis<br />

Calamagrostietum pseudophragmitis<br />

Salicetum elaeagni<br />

23<br />

Floodplain vegetation:<br />

Erico-Pinetum<br />

Alnetum incanae<br />

GPS-Coordinates of transect in investigation area 2 (from <strong>Workshop</strong> 20<strong>10</strong>):<br />

46°13`18.79”/13°1`21.64” (Altitude: 153 m)<br />

2%


Fig. 21: Location of the investigation area at Forni di Sotto (TK 25, Dolomiti di Friulane)<br />

4.4 FFH SITE IT33<strong>10</strong>001 <strong>–</strong> DOLOMITI FRIULANE<br />

Fig. 22: Delimitation of the FFH-Area Dolomiti Friulane (European Environmental Agency 2008)<br />

24


Territory size: 36740.0000 ha Biogeographic region: alpine<br />

Altitude maximum: 2706 m Altitude minimum: 450 m<br />

Tab. 2: European Union Habitats and their percentage of the area (European Environmental Agency)<br />

Code EUH Description<br />

25<br />

Percentage<br />

of area<br />

3220 <strong>Alpine</strong> rivers and the herbaceous vegetation along their banks 1%<br />

3240 <strong>Alpine</strong> rivers and their lingeous vegetation with Salix eleagnos 1%<br />

4060 <strong>Alpine</strong> and Boreal heaths 3%<br />

4070* Bushes with Pinus mugo and Rhododendron hirsutum (Mugo-<br />

Rhododendretum hirsuti)<br />

11%<br />

6170 <strong>Alpine</strong> and subalpine calcareous grasslands 12%<br />

62A0 Eastern sub-mediteranean dry grasslands (Scorzoneratalia villosae) 5%<br />

6430 Hydrophilous tall herb fringe communities of plains and of the montane to<br />

alpine levels<br />

1%<br />

8120<br />

Calcareous and calcshist screes of the montane to alpine levels (Thlaspietea<br />

rotundifolii)<br />

8160* Medio-European calcareous scree of hill and montane levels 3%<br />

82<strong>10</strong> Calcareous rocky slopes with chasmophytic vegetation 8%<br />

91K0 Illyrian Fagus sylvatica forests(Aremonio-Fagion) 28%<br />

94<strong>10</strong><br />

Acidophilous Picea forests of the montane to alpine levels (Vaccinio-<br />

Piceetea)<br />

9530* (Sub-) Mediterranean pine forests with endemic black pines 3%<br />

* Priority habitat<br />

8%<br />

4%


4.5 INVESTIGATION AREA 3 <strong>–</strong> AMARO / FELLA<br />

Fig. 23: Investigation area 3 <strong>–</strong> Amaro and alternative investigation area <strong>–</strong> Fella (TK 25, Dolomiti di Friulane)<br />

26


4.6 FFH SITE IT3320015 <strong>–</strong> VALLE DEL MEDIO TAGLIAMENTO<br />

Fig. 24: Delimitation of the FFH-Area Valle del Medio Tagliamento (European Environmental Agency 2008)<br />

Territory size: 3580.0000 ha Biogeographic region: alpine<br />

Altitude maximum: 1478 m Altitude minimum: 158 m<br />

Tab. 3: European Union Habitats and their percentage of area (European Environmental Agency)<br />

Code EUH Description<br />

Percentage<br />

of area<br />

3220 <strong>Alpine</strong> rivers and the herbaceous vegetation along their banks 3%<br />

3230 <strong>Alpine</strong> river and their lingeous vegetation with Myricaria germanica 5%<br />

3240 <strong>Alpine</strong> rivers and their lingeous vegetation with Salix elaeagnos 5%<br />

5130 Juniperus communis formations on heaths or calcareous grasslands 1%<br />

61<strong>10</strong>* Rupicolous calcareous or basophilic grasslands of the Alysso-Sedion albi 1%<br />

6170 <strong>Alpine</strong> and subalpine calcareous grasslands 15%<br />

62A0 Eastern sub-mediteraneandry grasslands (Scorzoneratalia villosae) 15%<br />

6430<br />

Hydrophilous tall herb fringe communities of plains of the montane to alpine<br />

levels<br />

8160* Medio-European calcareous scree of hill and montane levels 2%<br />

82<strong>10</strong> Calcareous rocky slopes with chasmophytic vegetation 2%<br />

9180* Tilio-Acerion forests of slopes, screes and ravins 5%<br />

91K0 Illyrian Fagus sylvatica forests (Aremonio-Fagion) 32%<br />

91L0 Illyrian oak-hornbeam forests (Erythronio-Carpinion) 1%<br />

* Priority habitat<br />

27<br />

1%


4.7 INVESTIGATION AREA 4 <strong>–</strong> CORNINO & MONTE DI RAGOGNA<br />

Fig. 25: Location of the investigation area Cornino & Mt. Ragogna, as well as Hotel Ai Glicini & Camp<br />

Ground Ai Pioppi (TK 25, Friuli Venezia Giulia)<br />

Short description of the investigation area 4 Cornino (from Lippert et al. 1995):<br />

At the transition from the Alps to the <strong>Alpine</strong> foreland a change in the river structure takes<br />

place. The slope of the running water and transportation force are reduced. Because of an<br />

increased sedimentation, a large gravel cone from Gemona to Casarsa grew in the course<br />

if the centuries. Around this area, the floodplain has a wideness of more than one<br />

kilometer and in the investigation area at Cornino it grows up two kilometer. The<br />

investigation area is located on a height of 160 m above sea level. This segment of the<br />

Tagliamento spreads into various arborising branches. Based on spring floods, various<br />

parts are free from vegetation.<br />

On account of those prevailing conditions (gravel and sandy river banks, frequent flood<br />

waters) pioneer and floodplain species settled this area.<br />

Pioneer vegetation:<br />

Epilobio-Scrophularietum caninae<br />

Echinochloa crus-gali-Xanthium<br />

strumaria-community<br />

Calamagrostietum pseudophragmitis<br />

Salici-Myricarietum<br />

28<br />

Salicetum elaeagni typicum<br />

Floodplain vegetation:<br />

Salici-Hippophaetum rhamnoidis<br />

Fraxinus ornus-Ostrya carpinifoliacommunity


GPS-Coordinates of transect in investigation area 4 (from <strong>Workshop</strong> 20<strong>10</strong>):<br />

East: 46°13`23.61”/13°1`22.48” (Altitude: 149 m)<br />

North: 46°13`23.61”/13°1`22.48” (Altitude: 149 m)<br />

South: 46°13`24.8”/13°1`24.21” (Altitude 162 m)<br />

Short description of the investigation area Monte di Ragogna (from <strong>Workshop</strong> 20<strong>10</strong>):<br />

This mountain (Altitude 512 m) lies directly to the floodplain of the Tagliamento and is<br />

located near the city Ragogna. From the top of the Monte di Ragogna, a huge and<br />

impressing view of the riverbed is possible. The large white banks and the several streams<br />

are plainly visible. On the east side of the mountain there is a pine forest and on the northwest<br />

side a forest with Ostrya carpinifolia got established.<br />

4.8 FFH SITE IT33<strong>10</strong>007 <strong>–</strong> GRETO DEL TAGLIAMENTO<br />

Fig. 26: Delimitation of the FFH-Area Greto del Tagliamento (European Environmental Agency, 2008)<br />

Territory size: 2719.0000 ha Biogeographic region: continental<br />

Altitude maximum: 164 m Altitude minimum: 92 m<br />

Tab. 4: European Union Habitats and their percentage of the area (European Environmental Agency)<br />

Code EUH Description<br />

29<br />

Percentage<br />

of area<br />

3220 <strong>Alpine</strong> rivers and the herbaceous vegetation along their banks 41%<br />

3240 <strong>Alpine</strong> rivers and their lingeous vegetation with Salix eleagnos 25%<br />

62A0 Eastern sub-mediteranean dry grasslands (Scorzoneratatlia villosae) <strong>10</strong>%


4.9 INVESTIGATION AREA 5 <strong>–</strong> SPILIMBERGO<br />

Fig. 27: Location of the investigation area at Spilimbergo (TK 25, Friuli Venezia Giulia)<br />

Short description of the investigation area 5 Spilimbergo (from Lippert et al. 1995):<br />

The investigation area has a height of <strong>10</strong>0 m above sea level. At this part, the Tagliamento<br />

has a wide of 3 km and consist of several small arms, which keep on separating and<br />

combining. Along the river arms, there are sand and gravel banks with different stages of<br />

floodplain succession. The sedimentation of sand and silt has risen, but at some parts<br />

have been rinsed, so that gravel banks are present.<br />

Pioneer vegetation:<br />

Epilobio-Scrophularietum caninae<br />

Echinochloa crus-gali-Xanthium<br />

strumaria-community<br />

Calamagrostietum pseudophragmitis<br />

Amorpha fruticosa-community<br />

Salicetum elaeagni typicum<br />

30<br />

Floodplain vegetation:<br />

Salicetum elaeagni typicum<br />

Amorpha fruticosa <strong>–</strong>community


4.<strong>10</strong> INVESTIGATION AREA 6 <strong>–</strong> CASARSA<br />

Fig. 28: Location of the investigation area at Casarsa (TK 25, Friuli Venezia Giulia)<br />

4.11 INVESTIGATION AREAS 7 & 8 <strong>–</strong> BOLZANO & LATISANA<br />

Fig. 29: Location of the investigation area at Bolzano and Latisana (TK 25, Friuli Venezia Giulia)<br />

31


4.12 INVESTIGATION AREA 9 <strong>–</strong> BIBIONE / ESTUARY<br />

Fig. 30: Location of the investigation area of the estuary at Bibione (TK 25, Friuli Venezia Giulia)<br />

4.13 FFH SITE IT3250033 <strong>–</strong> LAGUNA DI CAORLE / FOCE DEL TAGLIAMENTO<br />

Fig. 31: Delimitation of the FFH-Area Laguna di Caorle / Foce del Tagliamento (European Environmental<br />

Agency, 2012)<br />

Territory size: 4386 ha Biogeographic region: continental<br />

Altitude maximum: 2 m Altitude minimum: 0 m<br />

32


Tab. 5: European Union Habitats and their percentage of the area (European Environmental Agency)<br />

Code EUH Description<br />

33<br />

Percentage<br />

of area<br />

1150* Coastal lagoons 15%<br />

12<strong>10</strong> Annual vegetation of drift lines 1%<br />

13<strong>10</strong> Salicornia and other annuals colonizing mud and sand 4%<br />

1420<br />

Mediterranean and thermo-Atlantic halophilous scrubs (Sarcocornetea<br />

fruticosi)<br />

15<strong>10</strong>* Mediterranean salt steppes (Limonietalia) 16%<br />

21<strong>10</strong> Embryonic shifting dunes 1%<br />

2120 Shifting dunes along the shoreline with Ammophila arenaria ('white dunes') 1%<br />

2130* Fixed coastal dunes with herbaceous vegetation ('grey dunes') 2%<br />

2230 Malcolmietalia dune grasslands 1%<br />

2250* Coastal dunes with Juniperus spp 2%<br />

2270* Wooded dunes with Pinus pinea and/or Pinus pinaster 12%<br />

64<strong>10</strong><br />

Molinia meadows on calcareous, peaty or clayey-silt-laden soils (Molinion<br />

caeruleae)<br />

6420 Mediterranean tall humid grasslands of the Molinio-Holoschoenion 5%<br />

72<strong>10</strong>*<br />

Calcareous fens with Cladium mariscus and species of the Caricion<br />

davallianae<br />

9340 Quercus ilex and Quercus rotundifolia forests 5%<br />

* Priority habitat<br />

4.14 INVESTIGATION AREA <strong>10</strong> <strong>–</strong> FLAMBRO<br />

Fig. 32: Location of the investigation area at Flambro (TK 25, Friuli Venezia Giulia)<br />

3%<br />

1%<br />

1%


Short description of the investigation area <strong>10</strong> Flambro (from <strong>Workshop</strong> 20<strong>10</strong>):<br />

This FFH-Area is shaped by calcareous fens, which are fed by rain and spring water that<br />

comes out at the surface. There are a lot of endemic species settled at this location. This<br />

is an important sign for a long consistency of the site. Grasses are the dominating species<br />

for those low moors. The Life-Project established several hiking tracks in this relict<br />

location.<br />

4.15 FFH SITE IT3320026 <strong>–</strong> RISORGIVE DELLO STELLA<br />

Fig. 33: Delimitation of the FFH-Area Risorgive dello Stella (European Environmental Agency 2006)<br />

Territory size: 796.0000 ha Biogeographic region: continental<br />

Altitude maximum: 36 m Altitude minimum: 9 m<br />

Tab. 6: European Union Habitats and their percentage of the area (European Environmental Agency)<br />

Code EUH Description<br />

34<br />

Percentage<br />

of area<br />

3140 Hard oligo-mesotrophic waters with benthic vegetation of Chara spp. 1%<br />

3260 Water courses of plain to montane levels with the Ranunculion fluitans and<br />

Callitricho-Batrachion<br />

6%<br />

62A0 Eastern sub-mediteranean dry grasslands (Scorzoneratatlia villosae) 4%<br />

6430 Hydrophilous tall herb fringe communities of plains and of the montane to<br />

alpine levels<br />

1%<br />

64<strong>10</strong><br />

Molinia meadows on calcareous, peaty or clayey-silt-laden soils (Molinion<br />

caeruleae)<br />

6%<br />

72<strong>10</strong>* Calcareous fens with Cladium mariscus and species of the Caricion 4%<br />

7230 Alkaline fens 5%<br />

91E0* Alluvial forests with Alnus glutinosa and Fraxinus excelsior (Alno-Padion,<br />

Alnion incanae, Salicion albae)<br />

6%<br />

91F0<br />

*Priority habitat<br />

Riparian mixed forests of Quercus robur, Ulmus laevis, Ulmus minor,<br />

Fraxinus excelsior or Fraxinus angustifolia (Ulmenion minoris)<br />

8%


4.16 INVESTIGATION AREA 11 <strong>–</strong> VIVARO<br />

Fig. 34: Location of the investigation area at Vivaro (TK 25, Friuli Venezia Giulia)<br />

Short description of the investigation area 11 Vivaro (from <strong>Workshop</strong> 20<strong>10</strong>):<br />

At this area, in the past glacier flowed into the <strong>Alpine</strong> Foreland and formed a huge gravel<br />

body. The soil sustains of strong weathered sand-stone. The spatial leads temporarily<br />

water from the Alps.<br />

4.17 FFH SITE IT33<strong>10</strong>009 <strong>–</strong> MAGREDI DEL CELINA<br />

Fig. 35: Delimitation of the FFH-Area Magredi del Celina (European Environmental Agency 2007)<br />

35


Territory size: 4372.0000 ha Biogeographic region: continental<br />

Altitude maximum: 244 m Altitude minimum: 50 m<br />

Tab. 7: European Union Habitats and their percentage of the area (European Environmental Agency)<br />

Code EUH Description Percentage<br />

3220 <strong>Alpine</strong> rivers and the herbaceous vegetation along their banks 31%<br />

3240 <strong>Alpine</strong> rivers and their lingeous vegetation with Salix eleagnos 4%<br />

62A0 Eastern sub-mediteranean dry grasslands (Scorzoneratalia villosae) 53%<br />

36


5 METHODS OF RESEARCH<br />

5.1 ASSESSMENT OF VASCULAR PLANTS, KEY SPECIES & NEOPHYTES (NORBERT<br />

MÜLLER)<br />

Investigated will be all vascular plants, as well as certain key species and neophytes within<br />

the 11 investigation areas (see Fig. 18). The investigation areas 1-8 correspond to the first<br />

research workshops in 1991 & 1992 (Lippert et al. 1995).<br />

With a simple method the frequency of vascular plant species in alpine river landscapes<br />

will be recorded in repetition to the same investigations in 2004, 2005 and 2007.<br />

The status of neophytes is classified according to Poldini et al. (2002) and supplemented<br />

by Lohmeyer & Sukopp (1993) as well as Kowarik (2003).<br />

Approach:<br />

Transects were fixed in the respective investigation areas. They run straight through the<br />

floodplain and the riverbed (see Fig. 36). An area of approx. 50m right and left of transect<br />

is inspected in detail and in the fixed amount of time of two hours. The key species and<br />

neophytes are recorded by frequency in the<br />

a) Pioneer vegetation sites and<br />

b) Floodplain Forests<br />

The scale of frequency is<br />

1 <strong>–</strong> rare (only few individuals)<br />

2 <strong>–</strong> dispersed (occurrence with low frequency and/or low abundance)<br />

3 <strong>–</strong> common (occurrence with high frequency and/or high abundance)<br />

Fig. 36: River dynamics and structure of floodplain vegetation (simplified depiction) at the lower course of a<br />

debris-rich northern alpine river (alpine foreland) before river regulation (from Müller 1995).<br />

37


The following two tables contain key species and non-native species (neophytes) of<br />

vascular plants along the Tagliamento River. In addition, all occurring species are<br />

recorded in a flora list (see Tab. <strong>10</strong> in chapter 7).<br />

Tab. 8: Key species<br />

Investigation Areas<br />

1 2 3 4 5 6 7 8 9 <strong>10</strong> 11<br />

a = pioneer vegetation<br />

b = floodplain vegetation<br />

H = sample of species in herbarium of Tagliamento<br />

w orkshop<br />

No. H Species a b a b a b a b a b a b a b a b a b a b a b<br />

1 Aquilegia einseleana F.W.Schultz<br />

2 Barbarea vulgaris s.l. R.Br.<br />

3 Calamagrostis pseudophragmites (Haller f.) Koeler<br />

4 Campanula cespitosa Scop.<br />

5 Chondrilla chondrilloides (Ard.) H. Karst.<br />

6 Echium vulgare L.<br />

7 Epilobium dodonaei Vill.<br />

8 Galeopsis angustifolia Ehrh.<br />

9 Gypsophila repens L.<br />

<strong>10</strong> Hieracium piloselloides grex florentinum (All.) Zahn<br />

11 Hippophae rhamnoides subsp. fluviatilis v. Soest<br />

12 Leontodon berinii (Bartl.) Roth<br />

13 Linaria alpina (L.) Mill.<br />

14 Matthiola carnica Tammaro<br />

15 Melilotus alba L.W. Medicus<br />

16 Myricaria germanica (L.) Desv.<br />

17 Petasites paradoxus (Retz.) Baumg.<br />

18 Salix daphnoides Vill.<br />

19 Salix eleagnos Scop.<br />

20 Salix myrsinifolia Salisb. (= Salix nigricans )<br />

21 Scrophularia canina L. (incl. S. juratensis )<br />

22 Silene vulgaris subsp. glareosa (Jord.) M.-J. & Turrill<br />

Mauria<br />

38<br />

Forni di<br />

Sotto<br />

Amaro<br />

Cornino<br />

Spilimbergo<br />

Casarsa<br />

Tab. 9: Neophytes<br />

Investigation Areas 1 2 3 4 5 6 7 8 9 <strong>10</strong> 11<br />

a = pioneer vegetation<br />

b = floodplain vegetation<br />

H = sample of species in herbarium of Tagliamento<br />

w orkshop<br />

No. H Species a b a b a b a b a b a b a b a b a b a b a b<br />

1 Ailanthus altissima (Mill.) Sw ingle<br />

2 Ambrosia artemisifolia L.<br />

3 Amorpha fruticosa L.<br />

4 Bidens frondosa L.<br />

5 Buddleja davidii Franch.<br />

6 Conyza canadensis (L.) Cronq.<br />

7 Eragrostis pectinacea (Michx.) Nees<br />

8 Erigeron annus (L.) Pers.<br />

9 Galinsoga ciliata (Raf.) Blake<br />

<strong>10</strong> Helianthus tuberosus L.<br />

11 Impatiens glandulifera Royle<br />

12 Impatiens parviflora DC.<br />

13 Matricaria discoidea DC.<br />

14 Oenothera biennis agg.<br />

15 Panicum capillare L.<br />

16 Phalaris canariensis<br />

17 Populus x canadensis Moench<br />

18 Robinia pseudoacacia L.<br />

19 Senecio inaequidens DC.<br />

20 Solidago canadensis L.<br />

21 Solidago gigantea var. serotina (O. Kuntze) Cronq.<br />

22 Xanthium spec.<br />

Mauria<br />

Forni di<br />

Sotto<br />

Amaro<br />

Cornino<br />

Spilimbergo<br />

Casarsa<br />

Bolzano<br />

Bolzano<br />

Latisana<br />

Latisana<br />

Estuary<br />

Estuary<br />

Flambro<br />

Flambro<br />

Vivaro<br />

Vivaro


5.2 RESEARCH PROGRAMME ANIMAL ECOLOGY (NORBERT GROSSER)<br />

We continue recording animal fauna of Tagliamento area in the different investigation<br />

areas (see Fig. 18 & 36) from the last years in this short aspect of beginning summer.<br />

Investigation will comprise the following animal groups:<br />

- Butterflies and moths<br />

- Grasshoppers<br />

- Dragonflies<br />

- Amphibians and reptiles<br />

- Additionally snails, birds and mammals according to their biodiversity and existence of<br />

key species in river and adjacent habitats<br />

Because there is no identity between zoocoenoses, biotop types and vegetation units the<br />

investigation of fauna will be separated from the single habitat investigation and a<br />

combined transect method across all habitat-types in a defined river section will be used.<br />

Zoocoenoses are connected with habitats either by trophical relations between plants and<br />

animals, structural relations, microclimate in a habitat or successional stages of vegetation<br />

(indirectly), biotopnetwork and migration ability.<br />

Methods are:<br />

- Transect mapping (visual and acoustic observation)<br />

- Capture of insects with different nets and other methods (release after determination)<br />

- Observation of nocturnal active insects at light (in the surroundings of Cornino)<br />

- Exhaustor (in connection with investigation of soil substrates)<br />

39


6 POSTER SERIES "TAGLIAMENTO <strong>–</strong> KING OF ALPINE RIVERS"<br />

The posters on the following pages are results of the workshops in 2006 and 2009.<br />

Poster <strong>10</strong> of 2006 and Poster 4 of 2009 are not available.<br />

40


2006-1 <strong>International</strong> <strong>Alpine</strong> <strong>Workshop</strong> <strong>–</strong> Tagliamento 2006<br />

41


2006-2 Geology and geography of Tagliamento River<br />

42


2006-3 Investigation in plant diversity<br />

43


2006-4 Investigation of animal biodiversity<br />

44


2006-5 Herbaceous vegetation of alpine river banks on sand<br />

45


2006-6 Herbaceous vegetation of alpine river banks on gravel<br />

46


2006-7 Herbaceous vegetation of alpine rivers with Myricaria germanica<br />

47


2006-8 Ligneous vegetation of alpine rivers with Salix eleagnos<br />

48


2006-9 Muddy banks of rivers with Chenopodion rubri and Bidention<br />

49


2006-11 Alluvial forest with Alnus glutinosa and Fraxinus excelsior<br />

50


2006-12 Riparian mixed forest along the great rivers (Ulmenion minoris)<br />

51


2009-1 Occurring birds and their biotopes at Tagliamento<br />

52


2009-2 Dragonflies (Odonata) <strong>–</strong> The Tagliamento as habitat<br />

53


2009-3 Investigation of amphibian occurrence<br />

54


2009-5 Locust species <strong>–</strong> Habitats of sparse vegetation<br />

55


7 FLORA OF THE TAGLIAMENTO (NORBERT MÜLLER)<br />

Tab. <strong>10</strong>: Flora along the Tagliamento River<br />

Müller, N. (2006): Flora Tagliamento (Friaul, Italy) w ith key species and non-native species - Results from investigations 1991-1993, 2004<br />

and 2005. - University of Applied Sciences Erfurt: 12pages, n.p.<br />

Further involved researches: Schauer, Th., Rossel, S. & Lippert, W. (1991-1993), Bangert, U. & Drescher, A. (2004), von Heßberg, A. &<br />

Poldini, L. (2005), Abendroth, S. & Ramseier, D. (2006)<br />

Legend: X = Registered betw een 1991 and 20<strong>10</strong><br />

= Space for the registration 2012<br />

a = Pioneer vegetation<br />

b = Floodplain vegetation<br />

H= Sample of species in herbarium of Tagliamento w orkshop<br />

Investigation Areas: 1 2 3 4 5 6 7 8 9<br />

No. H Species a b a b a b a b a b a b a b a b a b a b a b<br />

1 Abies alba L. X . . . . . . .<br />

2 Acer campestre L. . . X X . . . .<br />

3 Acer negundo L. . . . . . . X .<br />

4 Acer pseudoplatanus L. X X X X . . . .<br />

5 Achillea millefolium L. . X X X X X . .<br />

6 Achnatherum calamagrostis (L.) PB. . . X X X X . .<br />

7 Acinos arvensis (Lam.) Dandy . . . X X . . .<br />

8 Aconitum lycoctonum L. (= A. vulparia Rchb. ex Spreng.) . X . . . . . .<br />

9 Adenostyles glabra (Mill.) DC. X . . . . . . .<br />

<strong>10</strong> Aegopodium podagraria L. . X X X . . X .<br />

11 Aethionema saxatile (L.) R.BR. . X . . . . . .<br />

12 Agropyron caninum (L.) PB. . X X X . . . .<br />

13 Agropyron pungens (Pers.) Roem. & Schult. . . X X X X X .<br />

14 Agropyron repens (L.) P.B. (oder in 7 bis 8 A. pungens ?) . . . . . X X X<br />

15 Agrostis capillaris L. subsp. capillaris . . . . . . . .<br />

16 Agrostis gigantea Roth . X X X X X X X<br />

17 Agrostis stolonifera L. . X X X X X X X<br />

18 Alchemilla vulgaris agg. . X . . . . . .<br />

19 Alisma lanceolatum With. . . X X . . . .<br />

20 Alisma plantago-aquatica L. . . . . . X X .<br />

21 Allium carinatum L. s.l. . . X X . . . .<br />

22 Alnus glutinosa (L.) Gaertn. . . . X X . X .<br />

23 Alnus incana (L.) Moench X X X X X X . .<br />

24 Ambrosia artemisiifolia L. . . . X X X X .<br />

25 Amelanchier ovalis Med. X X X X . . . .<br />

26 H Amorpha fruticosa L. . . . X X X X X<br />

27 Anagallis arvensis L. . . X X . X X .<br />

28 Anemone trifolia L. X X . . . . . .<br />

29 Angelica sylvestris L. . X X . X X . .<br />

30 H Anthyllis vulneraria L. s.l. . X . . . . . .<br />

31 Aposeris foetida (L.) Less. X X . . . . . .<br />

32 Aquilegia atrata Koch X . . . . . . .<br />

33 H Aquilegia einseleana F.W.Schultz (Endemit) X X X . . . . .<br />

34 Arabis pumila Jacq. subsp. stellulata (Bertol.) Nyman X . . . . . . .<br />

35 Arctium nemorosum Lej. . . . . . . X .<br />

36 Arctostaphylos uva-ursi (L.) Spreng. X X . . . . . .<br />

37 Arenaria serpyllifolia L. . . . . X . . .<br />

38 Arrhenatherum elatius (L.) J.&C.Presl. . . . . . . . X<br />

39 Artemisia alba Turra . . . . . . . .<br />

40 Artemisia absinthium L. . . X . . . . .<br />

41 Artemisia campestris L. . . X X X X X .<br />

42 Artemisia verlotiorum Lamotte . . . . . . X .<br />

43 Artemisia vulgaris L. . X X X X X X X<br />

44 Arum cf. maculatum L. . . . . . . X .<br />

45 Aruncus dioicus (Walter) Fernald X X X . . . . .<br />

46 Arundo donax L. . . . . . . X .<br />

Mauria<br />

56<br />

Forni di<br />

Sotto<br />

Amaro<br />

Cornino<br />

Spilimbergo<br />

Casarsa<br />

Bolzano<br />

Latisana<br />

Estuary<br />

<strong>10</strong> 11<br />

Flambro<br />

Vivaro


Investigation Areas: 1 2<br />

No. H Species a b a b a b a b a b a b a b a b a b a b a b<br />

47 Asparagus officinalis L. . . . X X X X .<br />

48 Asperula aristata (L.) L. f. subsp. scabra (Presl) Nym X X X . . . . .<br />

49 Asperula cynanchica L. . . X X X X X .<br />

50 Asperula purpurea (L.) Ehrend. . . . X X . . .<br />

51 Asplenium ruta-muraria L. . . . X . . . .<br />

52 Asplenium trichomanes L. . . . X . . . .<br />

53 Asplenium viride Huds. X . . . . . . .<br />

54 Aster bellidiastrum (L.) Scop. X X . . . . . .<br />

55 Aster novi-belgii L. (agg.) . . . . . . X .<br />

56 H Astragalus onobrychis L. . . X X X . X .<br />

57 Astragalus penduliflorus Lam. X . . . . . . .<br />

58 Astragalus cf. purpureus Lam. . . X X . . . .<br />

59 H Barbarea vulgaris R. Br. . X . . . . X .<br />

60 Bellis perennis L. . X . . . . . .<br />

61 Berberis vulgaris L. . X X X . . . .<br />

62 Berula erecta (Huds.) Coville (= Sium erectum Huds.) . . X X . . X .<br />

63 Betonica alopecurus L. . X . . . . . .<br />

64 Betonica hirsuta L. . X . . . . . .<br />

65 Betula pendula Roth X X . . . . . .<br />

66 Bidens cernua L. . . . . . X X .<br />

67 H Bidens frondosa L. . . X X X X X .<br />

68 H Bidens tripartita L. . . X X X X X .<br />

69 H Biscutella laevigata L. X X X . . . . .<br />

70<br />

Blackstonia acuminata (W.D.J.Koch & Ziz) Domin<br />

subsp. acuminata<br />

. . . . . . .<br />

71 H Blackstonia perfoliata (L.) Huds. subsp. perfoliata . X . . X X X .<br />

72 H Bothriochloa ischaemum (L.) Keng . . . . X X X .<br />

73 Brachypodium pinnatum agg. . . X . . . . .<br />

74 H Brachypodium rupestre (Host.) Roem. & Schult. X X X X X X . X<br />

75 Brachypodium sylvaticum (Huds.) PB. . X X X X X X .<br />

76 Brassica cf. nigra L. . . . . X . . .<br />

77 Briza media L. X X . . . . . .<br />

78 H Bromus erectus Huds. s.s. . X . X . X . .<br />

79 Bromus hordeaceus agg. . . . . . . X .<br />

80 Bromus sterilis L. . . . . . X . .<br />

81 Buddleja davidii Franch. . . X X X X X .<br />

82 Buphthalmum salicifolium L. X X X X X X X .<br />

83 Butomus umbellatus L. . . X X . . . .<br />

84 Calamagrostis arundinacea (L.) Roth . . . . X . . .<br />

85 Calamagrostis pseudophragmites (Hall. f.) Koel. . X X X X X X X<br />

86 Calamagrostis varia (Schrad.) Host X X X . X X . .<br />

87 Callitriche hermaphroditica L. . . . . . . X .<br />

88 Callitriche palustris agg. . . . . . . X .<br />

89 Calystegia sepium (L.) R.Br. . . X X X . X .<br />

90 Campanula cespitosa Scop. X X X X . . . .<br />

91 Campanula sibirica L. . . . . . X . .<br />

92 Campanula trachelium L. . X X X . . . .<br />

93 Capsella bursa-pastoris (L.) Med. . X . . X . . .<br />

94 Cardamine flexuosa With. . X . . . . . .<br />

95 Cardamine impatiens L. . . X . . . . .<br />

96 Cardamine amara L. subsp. amara . . X X . . X .<br />

97 Carduus defloratus L. sensu Poldini . X X X X X . .<br />

98 Carex cf. acutiformis Ehrh. . . X X . . . .<br />

99 Carex alba Scop. X X X X X . . .<br />

Carex brachystachys Schrank (1,2 statt C. ferruginea<br />

<strong>10</strong>0 H<br />

1995)<br />

X X . . . . . .<br />

<strong>10</strong>1 Carex caryophyllea Latourr. . X . . . . . .<br />

<strong>10</strong>2 Carex digitata L. . X X X . . . .<br />

<strong>10</strong>3 Carex echinata Murray X . . . . . . .<br />

<strong>10</strong>4 Carex elata All. . . X . . . . .<br />

<strong>10</strong>5 Carex ferruginea Scop. X . . . . . . .<br />

<strong>10</strong>6 Carex firma Host X . . . . . . .<br />

Mauria<br />

57<br />

Forni di<br />

Sotto<br />

3 4 5 6 7 8 9 <strong>10</strong> 11<br />

Amaro<br />

Cornino<br />

Spilimbergo<br />

Casarsa<br />

Bolzano<br />

Latisana<br />

Estuary<br />

Flambro<br />

Vivaro


Investigation Areas: 1 2<br />

No. H Species a b a b a b a b a b a b a b a b a b a b a b<br />

<strong>10</strong>7 Carex flacca Schreb. X X X X X X . .<br />

<strong>10</strong>8 H Carex flava (agg.) L. s.l. X X X X . . . .<br />

<strong>10</strong>9 Carex gracilis Curtis (= C. acuta L.) . . X . . . . .<br />

1<strong>10</strong> Carex hallerana Asso . . X X . . . .<br />

111 Carex hirta L. . . . . X . . .<br />

112 H Carex hostiana DC. . . X . . . . .<br />

113 Carex humilis Leyser . X X . . . . .<br />

114 Carex lepidocarpa Tausch (det. Podlech, in 1) X . . . . . . .<br />

115 Carex montana L. . X . . . . . .<br />

116 H Carex mucronata All. . X X X . X . .<br />

117 Carex oederi Retz. . X . . . . . .<br />

118 Carex ornithopoda Willd. X X . X . . . .<br />

119 Carex panicea L. X . X . . . . .<br />

120 Carlina acaulis L. . X . . . . . .<br />

121<br />

Carlina biebersteinii subsp. brevibracteata (Andrae)<br />

K.Werner<br />

. . . . X . . .<br />

122 Carlina vulgaris L. . X X X . X . .<br />

123 Centaurea dichroantha Kern. . . X X X X . .<br />

124 H Centaurea jacea L. . X X X X X X .<br />

125 Centaurea forojulensis (Poldini) Poldini . . . . X . X .<br />

126 Centaurea maculosa Lam. . . X X X X . .<br />

127 Centaurea nemoralis Jordan . . . . . X . .<br />

128<br />

Centaurea nigrescens Willd. subsp. transalpina (Schl.)<br />

Nyman<br />

X X . . X X . .<br />

129<br />

Centaurea nigrescens Willd. subsp. vochinensis (Bernh.<br />

ex Rchb.) Nyman<br />

. X . . X . . .<br />

130 Centaurea rhenana Bor. (= C. stoebe L.) . . . X . X X X<br />

131 H Centaurea scabiosa L. . . . X . . . .<br />

132 Centaurea scabiosa L. subsp. fritschii (Hayek) Hayek . . X X . X . .<br />

133 Centaurea scabiosa L. subsp. scabiosa . X . . . . . .<br />

134 Centaurium erythraea Raf n subsp. erythraea . . X . X X X .<br />

135<br />

Centaurium majus (H. et L.) Zeltner (Prot. Müller 1992,<br />

S.29)<br />

. . . X . . . .<br />

136 Centaurium pulchellum (Sw.) Druce . . . . . X X .<br />

137 Cephalanthera damasonium (Mill.) Druce X . . . . . . .<br />

138 Cerastium alpinum L. . X . . . . . .<br />

139 H Cerastium carinthiacum Vest subsp. carinthiacum X X . . . . . .<br />

140 Cerastium holosteoides Fries em. Hyl. . X . . . . . .<br />

141 Chaenorhinum minus (L.) Lange subsp. minus . X X X X X X .<br />

142 Chaerophyllum hirsutum L. . X . . . . . .<br />

143 Chamaecytisus hirsutus (L.) Link. X X . . . . . .<br />

144 Chamaecytisus purpureus (Scop.) Lk. X X X . . . . .<br />

145 Chenopodium album L. . X X X X X X .<br />

146 Chenopodium rubrum L. . . X . . . . .<br />

147 H Chondrilla chondrilloides (Ard.) Karsten . X X X . . . .<br />

148 Cichorium intybus L. . . X X X X X .<br />

149 Circea intermedia Ehrh. . . X . . . . .<br />

150 Cirsium arvense (L.) Scop. . . X X . X X X<br />

151 H Cirsium erisithales (Jacq.) Scop. X X . . . . . .<br />

152 Cirsium heterophyllum (L.) Hill X . . . . . . .<br />

153 Cirsium oleraceum (L.) Scop. . . X X . . . .<br />

154 Cirsium vulgare (Savi) Ten. . . X X . . X .<br />

155 Clematis alpina Mill. X . . . . . . .<br />

156 Clematis recta L. . X . . X . . .<br />

157 Clematis vitalba L. . X X X X X X .<br />

158 Convallaria majalis L. X X X X X . . .<br />

159 Convolvulus arvensis L. . . X . . . . .<br />

160 Conyza canadensis (L.) Cronq. . X X X X X X .<br />

161 Cornus sanguinea L. X X X X X X X .<br />

162 Cornus sanguinea subsp. hungarica (Kárpáti) Soó . . . . . X X .<br />

163 Coronilla emerus L. subsp. emerus . . . X . . . .<br />

164 Coronilla vaginalis Lam. . X . . . . . .<br />

Mauria<br />

58<br />

Forni di<br />

Sotto<br />

3 4 5 6 7 8 9 <strong>10</strong> 11<br />

Amaro<br />

Cornino<br />

Spilimbergo<br />

Casarsa<br />

Bolzano<br />

Latisana<br />

Estuary<br />

Flambro<br />

Vivaro


Investigation Areas: 1 2<br />

No. H Species a b a b a b a b a b a b a b a b a b a b a b<br />

165 Corylus avellana L. X . X X . X X .<br />

166 Cotoneaster integerrimus Med. . X . . . . . .<br />

167 Crataegus monogyna (agg.) . X X X X . X .<br />

168 Crepis foetida L. (in 5 subsp. rhoedifolia ?) . . . . X X . .<br />

169 H Crepis rhoeadifolia M.Bieb. . . . . X X . .<br />

170 H Crepis setosa Haller fil. . . . . X X . .<br />

171 Cruciata glabra (L.) Ehrend. . X . . . . . .<br />

172 Cruciata laevipes Opiz X X . . . . . .<br />

173 Cuscuta campestris Yuncker . . . . . X . .<br />

174 Cuscuta epithymum (L.) L. . X . . . . . .<br />

175 Cyclamen purpurascens Mill. X X X . . . . .<br />

176 H Cynodon dactyolon (L.) Pers. . . . . . X . .<br />

177 H Cyperus fuscus L. . . . . . . X .<br />

178 H Cyperus glomeratus L. . . . . . . X .<br />

179 Cytisus decumbens (Durande) Spach . X . . . . . .<br />

180 Cytisus nigricans L. s.l. . X . . . . . .<br />

181 H Cytisus pseudoprocumbens Markgr. . X . . . . . .<br />

182 H Dactylis glomerata L. s.l. X X X X X X X .<br />

183 Dactylorhiza maculata (L.) Soo X X . . . . . .<br />

184 Daphne mezereum L. X X . . . . . .<br />

185 Daphne striata Tratt. X X X X X X . .<br />

186 Daucus carota L. X X X X X X X X<br />

187 Deschampsia cespitosa (L.) PB. X X X X X X X .<br />

188<br />

Dianthus sternbergii Sieber ex Capelli (= D.<br />

monspessulanus subsp. waldsteinii (Sternb.) Nyman)<br />

. . X . . . . .<br />

189 H Dianthus sylvestris Wulf. subsp. sylvestris X . . . . . . .<br />

190 H Diplotaxis tenuifolia (L.) DC. . X X X X X X .<br />

191 H Dorycnium herbaceum Vill. . X . . . . . .<br />

192<br />

Dorycnium pentaphyllum Scop. subsp. germanicum<br />

(Gremli) Gams<br />

. . X X X X . .<br />

193 Dryas octopetala L. X X X . . . . .<br />

194 Dryopteris filix-mas (L.) Schott . X X X . . . .<br />

195 Duchesnea indica (Andrews) Focke . . . . . . X .<br />

196 H Echinochloa crus-galli (L.) PB. . . . X X X X .<br />

197 Echium vulgare L. . X X X X X X .<br />

198 H Eleocharis palustris L. . . X . . . X .<br />

199 Elodea canadensis Michx. . . . . . . X .<br />

200 Elytrigia intermedia (Host) Nevski . . X X . . . .<br />

201 Epilobium cullinum C.G.Gmel. . X X X . . . .<br />

202 H Epilobium dodonaei Vill. . X X X X X X .<br />

203 Epilobium hirsutum L. . X . . . . X .<br />

204 Epilobium montanum L. . X . . . . . .<br />

205 Epilobium parviflorum Schreber . X . . . . . .<br />

206 Epilobium tetragonum L. . X . . . . . .<br />

207 Epipactis atrorubens (Hoffm.) Schult. X X . . . . . .<br />

208 Epipactis helleborine (L.) Crantz . X X . . . . .<br />

209 Epipactis palustris (L.) Cr. . X . . . . . .<br />

2<strong>10</strong> Equisetum arvense L. X X X X X X X X<br />

211 Equisetum fluviatile L. . X . . . . X .<br />

212 Equisetum hyemale L. . . X X . . X .<br />

213 Equisetum x meridionale (Milde) Chiov. . . . . . . X .<br />

214 H Equisetum palustre L. . X X . . . X .<br />

215 Equisetum palustre x variegatum Schleich. . X X X . . X .<br />

216 Equisetum ramosissimum Desf. . . . . . . X .<br />

217 Equisetum telmateja Ehrh. . X . . . . . .<br />

218 H Equisetum variegatum Schleich. ex Web. & Mohr . X X X . . X .<br />

219 H Eragrostis pectinacea (Michx.) Nees . . X X X X X .<br />

220 Erica carnea L. (= E. herbacea L.) X X X . . . . .<br />

221 Erica carnea L. subsp. carnea . X . . . . . .<br />

222 Erigeron annuus (L.) Pers. . X X X X X X X<br />

223 H Erigeron annuus (L.) Pers. subsp. annuus . X X X X . X .<br />

Mauria<br />

59<br />

Forni di<br />

Sotto<br />

3 4 5 6 7 8 9 <strong>10</strong> 11<br />

Amaro<br />

Cornino<br />

Spilimbergo<br />

Casarsa<br />

Bolzano<br />

Latisana<br />

Estuary<br />

Flambro<br />

Vivaro


Investigation Areas: 1 2<br />

Mauria<br />

No. H Species a b a b a b a b a b a b a b a b a b a b a b<br />

Erigeron annuus subsp. septendrionalis (Fernald<br />

224 H<br />

& Wiegand) Wagenitz<br />

. . . X X X . .<br />

225 Erigeron glabratus Hoppe . X . . . . . .<br />

226 Eriophorum angustifolium Honck. X . . . . . . .<br />

227 Erucastrum gallicum O.E.Schulz . . X . . . . .<br />

228 Euonymus europaea L. . . . X . . . .<br />

229 Eupatorium cannabinum L. . X X X X X X X<br />

230 Euphorbia amygdaloides L. . X . . . . . .<br />

231 Euphorbia cyparissias L. . X X X X X X .<br />

232 Euphorbia dulcis L. . X . . . . . .<br />

233 H Euphorbia falcata L. . . . X X X X .<br />

234 Euphorbia helioscopia L. . . . . . . . .<br />

235 Euphorbia nurans Lag. . . . . X . . .<br />

Euphorbia triflora subsp. kerneri (Huter) Poldini (= E.<br />

236 H<br />

kerneri Huter)<br />

X X X X X . . .<br />

237 H Euphrasia cuspidata Host X X . . . . . .<br />

238<br />

Euphrasia nemorosa (Pers.) Wallr. (ob E. stricta D.Wolf f<br />

.<br />

ex J.F.Lehm.?)<br />

X . . . . . .<br />

239 Euphrasia salisburgensis Funck ex Hoppe X X . . . . . .<br />

240 Euphrasia stricta D.Wolff ex J.F.Lehm. . X . . . . . .<br />

241 Fagus sylvatica L. X . X X . . . .<br />

242 Fallopia convolvolus (L.) A.Loeve. . . X X X . . .<br />

243 Festuca arundinacea Schreb. . X . X X X X .<br />

244 Festuca gigantea (L.)Vill. . . X X . . . .<br />

245 H Festuca norica (Hackel) Richter . . . X X X . .<br />

246 Ficus carica L. . X . X . . . .<br />

247 Filipendula ulmaria (L.) Maxim. subsp. ulmaria . X X X . . . .<br />

248 Fragaria vesca L. . X . . . . . .<br />

249 Frangula alnus Mill. X X X X X X X .<br />

250 Fraxinus excelsior L. . X X X X . . .<br />

251 H Fraxinus ornus L. X . X X X . . .<br />

252 Fumana procumbens Gren. & Godr. . . X X X X . .<br />

253 Galeopsis angustifolia (Ehrh.) Hoffm. . . X X X X X .<br />

254 Galeopsis pubescens Bess. . . . . . X . .<br />

255 Galeopsis speciosa Mill. . X X X . X . .<br />

256 Galeopsis tetrahit L. . . . X . . . .<br />

257 Galinsoga ciliata (Rafin.) Blake . X X X . X X .<br />

258 Galinsoga parviflora Cav. . . . X . . X .<br />

259 Galium anisophyllum Vill. X X . . . . . .<br />

260 Galium aparine L. . . X X . . X .<br />

261 Galium boreale L. . . . . . . . .<br />

262 H Galium laevigatum L. . . X . . . . .<br />

263 H Galium lucidum All. . X . X . X . .<br />

264 Galium megalospermum All. (= G.helveticum Weigel) . X . . . . . .<br />

265 Galium mollugo subsp. album (Mill.) Tzvelev X X X X X X X .<br />

266 Galium mollugo agg. . X X X . . . .<br />

267 H Galium palustre L. . . . . . X . .<br />

268 Galium sylvaticum L. X . X . . . . .<br />

269 Galium verum L. . X X X X X . .<br />

270 H Genista germanica L. . X X . . . . .<br />

271<br />

Genista radiata (L.) Scop. (= Cytisanthus radiatus (L.)<br />

O.F.Lang)<br />

. X . . . . . .<br />

272 Genista tinctoria L. . X . . . X . .<br />

273 Gentiana asclepiadea L. X . . . . . . .<br />

274 Gentianta clusii Perr. et Song. X X . . . . . .<br />

275 Gentianta utriculosa L. . X . . . . . .<br />

276 Geranium robertianum L. . X X X . . . .<br />

277 Geranium sanguineum L. . . X . . . . .<br />

278 Geranium sylvaticum L. X . . . . . . .<br />

279 Glechoma hederacea L. . X X . . . . .<br />

280 Globularia cordifolia L. X X X X . . . .<br />

281 Globularia punctata Lapeyr. . X . X X X . .<br />

60<br />

Forni di<br />

Sotto<br />

3 4 5 6 7 8 9 <strong>10</strong> 11<br />

Amaro<br />

Cornino<br />

Spilimbergo<br />

Casarsa<br />

Bolzano<br />

Latisana<br />

Estuary<br />

Flambro<br />

Vivaro


Investigation Areas: 1 2<br />

No. H Species a b a b a b a b a b a b a b a b a b a b a b<br />

282 Gymnadenia conopsea (L.) R.Br. . X . . . . . .<br />

283 H Gymnadenia odoratissima (L.) Rich. X X . . . . . .<br />

284 H Gymnocarpium robertianum (Hoffm.) Newman X . . . . . . .<br />

285 Gypsophila repens L. X X X X X X X .<br />

286 Hedera helix L. . . X X . . X .<br />

287 Helianthemum nummularium (L.) Mill. s.l. (agg.) . X X X . X X .<br />

288 Helianthemum ovatum (Viv.) Dunal . . . X X X . .<br />

289 Helianthus annuus L. . . . . . X . .<br />

290 Heli a nthus tuberosus L. . X X X X X X X<br />

291 Hemerocallis flava L. X . X . . . . .<br />

292 Heracleum sphondylium L. . . X X . . . .<br />

293 Hieracium bifidum Kit. ex Hornem. X . . . . . . .<br />

294 Hieracium cf. dollineri Schultz Bip. subsp. furcatum . . X . . . . .<br />

295 Hieracium murorum L. (= H. sylvaticum (L.) L.) X . . . . . . .<br />

Hieracium piloselloides Vill. grex florentinum<br />

296 H<br />

(All.) Zahn<br />

. X X X X X X .<br />

297 Hieracium piloselloides Vill. subsp. cylindriceps N. & P. . X . . . . . .<br />

298 H Hieracium porrifolium L. subsp. porrifolium X X . . . . . .<br />

299 H Hirochloë australis (Schrader) R. et Sch. . X . . . . . .<br />

300 Hippocrepis comosa L. . X . . X X . .<br />

301 Hippophaë rhamnoides subsp. fluviatilis v.Soest . . X X X X X .<br />

302 Holcus lanatus L. . . . . X . X X<br />

303 H Horminum pyrenaicum L. X X . . . . . .<br />

304 Humulus lupulus L. . . X X . . X .<br />

305 Hypericum montanum L. . X . . . . . .<br />

306 Hypericum perforatum L. . X X X X X X X<br />

307 Ilex aquifolium L. . . . X . . . .<br />

308 Impatiens balfourii Hook. f. . . X X X . X .<br />

309 Impatiens glandulifera Royle . X X X . . . .<br />

3<strong>10</strong> Impatiens noli-tangere L. . X . . . . . .<br />

311 Impatiens parviflora DC. . . X X . . . .<br />

312 H Inula ensifolia L. . . X . . . . .<br />

313 Iris pseudacorus L. . . . . . . X .<br />

314 Juglans regia L. . . X X . . . .<br />

315 Juncus alpino-articulatus Chaix X X X X X X . .<br />

316 H Juncus articulatus L. . X X X X . X .<br />

317 H Juncus bufonius L. . . X X X . X .<br />

318 Juncus conglomeratus L. . . X X X . . .<br />

319 Juncus inflexus L. . X X X . . . .<br />

320 Juncus ranarius Perr. & Song. . . X . . . . .<br />

321 Juncus subnodulosus Schrank . . X . . . X .<br />

322 Juncus tenuis Willd. . . X . . . . .<br />

323 Juniperus communis L. X X X X X . . .<br />

324 Kernera saxatilis (L.) Rchb. X X . . . . . .<br />

325 Knautia dipsacifolia Kreutz. X X X . . . . .<br />

326 Knautia cf. longifolia (W. & K.) Koch . X . . . . . .<br />

327 H Knautia ressmannii (Pacher) Brick. X X . . . . . .<br />

328 Koeleria gracilis Pers. . . . . . . . .<br />

329 Koeleria pyramidata (Lam.) PB. . X X X X X X .<br />

330 H Koeleria cf. splendens K.Presl . . . . X . . .<br />

331 Laburnum anagyroides Med. X X X . X . .<br />

332<br />

Lamiastrum flavidum (F. Herm.) Ehrend. (= Lamium<br />

galeobdolon (L.) L.)<br />

. X . . . . .<br />

333 Lamium montanum (Pers.) Hoffm. ex Kabath . X X . . . . .<br />

334 Lamium orvala L. . X . . . . .<br />

335 Larix decidua Mill. X X . . . . . .<br />

336 H Laserpitium peucedanoides L. X X . . . . . .<br />

337 H Laserpitium siler L. . X . X . . . .<br />

338 Lathyrus heterophyllus L. . X . . . . . .<br />

Lathyrus laevigatus subsp. occidentalis (Fischer &<br />

339 H<br />

Meyer) Breitstr.<br />

X . . . . . . .<br />

340 Lathyrus pratensis L. . X . . . . . .<br />

Mauria<br />

61<br />

Forni di<br />

Sotto<br />

3 4 5 6 7 8 9 <strong>10</strong> 11<br />

Amaro<br />

Cornino<br />

Spilimbergo<br />

Casarsa<br />

Bolzano<br />

Latisana<br />

Estuary<br />

Flambro<br />

Vivaro


Investigation Areas: 1 2<br />

No. H Species a b a b a b a b a b a b a b a b a b a b a b<br />

341 Lathyrus sylvestris L. . X . . . . . X<br />

342 Lathyrus vernus (L.) Bernh. . . X . . . . .<br />

343 H Leontodon berinii (Bartl.) Roth X X X X X X . .<br />

344 Leontodon hispidus L. s.l. X . X X X X . .<br />

Leontodon hispidus subsp. hyoseroides (Welw . ex<br />

345 H<br />

Rchb.) J.Murr<br />

. . . . X . . .<br />

346 Leontodon hispidus subsp. glabratus (W.D.J.Koch) . . . . X . . .<br />

347 H Leontodon incanus (L.)Schrank . . X X . . . .<br />

348 Leucanthemum heterophyllum (Willd.) DC. X X . . . . . .<br />

349 Leucanthemum vulgare Lam. . X . X X X . .<br />

350 Ligustrum vulgare L. . X X X X X X .<br />

351 Lilium martagon L. X . . . . . . .<br />

352 Linaria alpina (L.) Mill. . X X X X . . .<br />

353 Linum catharticum L. X X X . . . . .<br />

354 Linum tenuifolium L. . . . X X X X .<br />

355 Linum viscosum L. X . . . . . . .<br />

356 Lolium perenne L. . X X X . . X X<br />

357 Lomelosia graminifolia (L.) Greuter & Burnet . . . . . . . .<br />

358 Lonicera alpigena L. X X . . . . . .<br />

359 Lonicera japonica Thunb. . . . . . . X .<br />

360 Lonicera periclymenum L. . . X X . . . .<br />

361 Lonicera xylosteum L. . X X X . . . .<br />

362 Lotus corniculatus L. X X X X X X X .<br />

363 Lotus corniculatus subsp. hirsutus (W.D.J.Koch) Rothm. . X . . . . . .<br />

364 Lotus pilosus Jord. . . . . . . . .<br />

365 Luzula nivea (L.) DC. X . . . . . . .<br />

366 Lycopersicon esculentum Mill. . . . X . . . .<br />

367 Lycopodium annotinum L. X . . . . . . .<br />

368 Lycopus europaeus L. . . X X X X X .<br />

369 Lysimachia vulgaris L. . X X X X X X .<br />

370 Lythrum salicaria L. . . X X X X X .<br />

371 Maianthemum bifolium (L.) F.W.Schmidt . X . . . . . .<br />

372 Matricaria chamomilla L. . . X X . . . .<br />

373 M atricaria discoidea DC. . X X X . . . .<br />

374 M atthiola carnica Tammaro . . X X X X . .<br />

375 Medicago lupulina L. . X X X X X X .<br />

376 Medicago x varia Martyn . . . . . . X X<br />

377 Melampyrum sylvaticum L. subsp. sylvaticum X X . . . . . .<br />

378 Melica nutans L. X X X X . X . .<br />

379 M elilotus alba Med. . X X X X X X .<br />

380 Mentha aquatica L. . . X X . . X .<br />

381 Mentha longifolia (L.) Huds. emend. Harley . X X X X X X .<br />

382 Mentha cf. x rotundifolia (L.) Huds. . . . . X . . .<br />

383 Micromeria thymifolia (Scop.) Fritsch . . . . . . . .<br />

384 Milium effusum L. . . X . . . . .<br />

385 Moehringia ciliata (Scop.) DT. X . . . . . . .<br />

Molinia caerulea subsp. arundinacea (Schrank)<br />

386 H<br />

H.K.G.Paul<br />

X X X X X X X .<br />

387 Molinia caerulea (L.) Moench subsp. caerulea X X X X . . . .<br />

388 Mycelis muralis (L.) Dumort. . X . . . . . .<br />

389 Myosotis scorpioides L. . . X . . . X .<br />

390 Myosoton aquaticum (L.) Moench . X . . . . X .<br />

391 H Myricaria germanica (L.) Desv. . X X X . . . .<br />

392 Myriophyllum verticillatum L. . . . . X . . .<br />

393 Nasturtium officinale Ait. fil. . . X . . . . .<br />

394 Oenothera biennis L. agg. . X X X X X X .<br />

395 Oenothera oakesiana (A.Gray) Robbins . . . . X X . .<br />

396 Oenothera parviflora L. agg. . . . . . X X .<br />

397 Onobrychis viciifolia Scop. . . . . X . . .<br />

398 Ononis spinosa L. . X X . . X . .<br />

399<br />

Ophrys holosericea (Burm. fil.) Greuter (= O. fuciflora<br />

(F.W.Schmidt) Moench)<br />

. . . X . . . .<br />

Mauria<br />

62<br />

Forni di<br />

Sotto<br />

3 4 5 6 7 8 9 <strong>10</strong> 11<br />

Amaro<br />

Cornino<br />

Spilimbergo<br />

Casarsa<br />

Bolzano<br />

Latisana<br />

Estuary<br />

Flambro<br />

Vivaro


Investigation Areas: 1 2<br />

No. H Species a b a b a b a b a b a b a b a b a b a b a b<br />

400 Ophrys insectifera L. X . . . . . . .<br />

401 Origanum vulgare L. s.l. . X . . . . . .<br />

402 Orobanche gracilis Sm. X . . . . . . .<br />

403 Orthilia secunda (L.) House . X . . . . . .<br />

404 Ostrya carpinifolia Scop. . X X X X X X .<br />

405 Oxalis acetosella L. . X . . . . . .<br />

406 Oxalis corniculata L. . . . X . . X .<br />

407 Oxalis fontana Bunge . . . . . . X .<br />

Panicum capillare L. (bei 7 Protokoll Müller 1992,<br />

408 H<br />

S.51)<br />

. . . . X X X .<br />

409 Parietaria officinalis L. . . . . . . X .<br />

4<strong>10</strong> Paris quadrifolia L. . X . . . . . .<br />

411 Parnassia palustris L. X X . . . . . .<br />

412 Pastinaca sativa L. . X . X X . . .<br />

413 Pedicularis elongata A.Kerner X . . . . . . .<br />

414 Petasites hybridus (L.) G.,M. & Sch. . X X X . . X .<br />

415 Petasites paradoxus (Retz.) Baumg. X X X X X . . .<br />

416 H Petrorhagia saxifraga (L.) Lk. . . X X X X X .<br />

417 Peucedanum cervaria (L.) Lapeyr. . X . . . X . .<br />

418 Peucedanum oreoselinum (L.) Moench X . . X . X . .<br />

419 Peucedanum ostruthium (L.) Koch . X . . . . . .<br />

420 H Peucedanum verticillare (L.) Mert. & Koch. . X X X X X X .<br />

421 Phalaris arundinacea L. . . X X X X X .<br />

422 Phalaris canariensis L. . . X X . . . .<br />

423<br />

Phleum bertolonii DC. (= P. pratense subsp. bertolonii<br />

(DC.) Bornm.)<br />

. . X X . . . .<br />

424 Phleum pratense L. . X X X . . . X<br />

425<br />

Phragmites australis (Lav.)Trin. ex Steud. (= P.<br />

communis Trin.)<br />

. X X X . . X X<br />

426 Phyteuma orbiculare L. X . . . . . . .<br />

427 Picea abies (L.) Karsten X X X X . . . .<br />

428 Picris hieracioides L. . . X X . X X .<br />

429 Pimpinella saxifraga L. . . . . . X . .<br />

430 Pinus mugo Turra agg. X X . . . . . .<br />

431 Pinus nigra Arnold . . X X . . . .<br />

432 Pinus sylvestris L. X X X X . . . .<br />

433 Plantago lanceolata L. . . X X X X X X<br />

434<br />

Plantago major subsp. intermedia (Gilib.) Lange. (= P.<br />

uliginosa F.W.Schmidt, = P. intermedia DC.)<br />

. X . X X X X .<br />

435 Plantago major L. . X X X X X X .<br />

436 Plantago media L. . X . . . . . .<br />

437 Plantago serpentina All. (= P. strictissima L.) . . . . . X . .<br />

438 H Platanus x hybrida Brot. . . . . . . X .<br />

439 Platanus cf. orientalis L. . . . . . . . X<br />

440 Poa alpina L. X X . . . . . .<br />

441 Poa annua L. . X X X X . X .<br />

442 Poa compressa L. . X X X X X X .<br />

443 Poa palustris L. . . X X . . X .<br />

444 Poa pratensis L. . . X X . X . .<br />

445 Poa trivialis L. . X X X X . X .<br />

446 Polygala armara L. agg. . X . . . . . .<br />

447 Polygala chamaebuxus L. X X . . . . . .<br />

448 Polygala comosa Schkuhr X X X . . . . .<br />

449 Polygala nicaeensis subsp. forojulensis (A.Kern.) . X . . . . . .<br />

450 Polygonatum verticillatum (L.) All. X . . . . . . .<br />

451 Polygonum aviculare L. agg. . X X X . X X .<br />

452 Polygonum hydropiper L. . . . . . . X .<br />

453 H Polygonum lapathifolium L. . X X X X X X .<br />

454 Polygonum persicaria L. . X X X X X X .<br />

455 Polygonum viviparum L. X . . . . . . .<br />

456 Populus alba L. . . . . . . X .<br />

457 Populus x canescens (Ait.) Sm. X X X X . . . .<br />

Mauria<br />

63<br />

Forni di<br />

Sotto<br />

3 4 5 6 7 8 9 <strong>10</strong> 11<br />

Amaro<br />

Cornino<br />

Spilimbergo<br />

Casarsa<br />

Bolzano<br />

Latisana<br />

Estuary<br />

Flambro<br />

Vivaro


Investigation Areas: 1 2<br />

No. H Species a b a b a b a b a b a b a b a b a b a b a b<br />

458 H Populus nigra L. (incl. P. x canadensis Moench) . . X X X X X X<br />

459 Populus tremula L. X . . . . . . .<br />

460 Potamogeton natans L. . . . . . . X .<br />

461 Potamogeton perfoliatus L. . . X X X . X .<br />

462 Potentilla erecta (L.) Raeuschel X X . . . . . .<br />

463 Potentilla reptans L. . . . X . . . .<br />

464 Primula vulgaris Huds. (= P. acaulis (L.) Hill) . . X X . . . .<br />

465 Prunella grandiflora (L.) Scholler X X . X X . X .<br />

466 Prunella vulgaris L. . . X X X . . .<br />

467 Prunus avium L. . X X X . . . .<br />

468 Prunus mahaleb L. . . . X . . . .<br />

469 Pulicaria dysenterica (L.) Bernh. . . . . . X X .<br />

470 Pulmonaria mollis agg. . X . . . . . .<br />

471 Pyrola minor L. X . . . . . . .<br />

472 Pyrola rotundifolia L. X . . . . . . .<br />

473 H Quercus pubescens Willd. . . . X X . . .<br />

474 Ranunculus acris L. . X X X . . . .<br />

475 Ranunculus aquatilis L. . . . X . . . .<br />

476 Ranunculus circinatus Sibth. . . . x . . . .<br />

477 Ranunculus lanuginosus L. . X . . . . . .<br />

478 Ranunculus nemorosus DC. (3: s.str.) . X X . . . . .<br />

479 Ranunculus repens L. . X X X X . . .<br />

480 Reseda lutea L. . X X X X X X .<br />

481 Rhamnus saxatilis Jacq. . X X . X . . .<br />

482 Rhamnus catharticus L. . . X X . . . .<br />

483<br />

Rhinanthus aristatus Celak. (Achtung! ist eventuell der<br />

Endemit Rh. pampanini Chabert)<br />

X X . X X X . .<br />

484 Rhinanthus freynii (Kerner) Fiori . X . . . . . .<br />

485 Rhododendron hirsutum L. X X . . . . . .<br />

486 Ribes nigrum L. . X . . . . . .<br />

487 Robinia pseudoacacia L. . . X X X . X .<br />

488 Rorippa palustris (L.) Bess. emend. Jons. . . . . . . X .<br />

489 Rosa canina L. . . . . . . . .<br />

490 Rosa glauca Pourr. non Vill. . . X . . . . .<br />

491 Rosa pendulina L. X X . . . . . .<br />

492 Rubus caesius L. . X X X X X X X<br />

493 Rubus corylifolius agg. . . . . . . X .<br />

494 Rubus fruticosus agg. . X . X . . . .<br />

495 Rubus idaeus L. . X . . . . . .<br />

496 Rubus saxatilis L. X X . . . . . .<br />

497 H Rubus ulmifolius Schott s.l. . . . . X . X .<br />

498 Rumex conglomeratus Murray . . . . . X . .<br />

499 Rumex crispus L. . X . . X . . .<br />

500<br />

Sacharum ravenae (L.) Murray (= Erianthus ravennae<br />

(L.) PB.)<br />

. . . . . . X .<br />

501 Salix alba L. . X X X X X X .<br />

502 Salix appendiculata Vill. X X X . . . . .<br />

503 Salix caprea L. . X . . . . . .<br />

504 H Salix daphnoides Vill. X X X X X X . .<br />

505 Salix daphnoides Vill. x nigricans Sm. . X . . . . . .<br />

506 H Salix elaeagnos Scop. X X X X X X X .<br />

507 H Salix glabra Scop. X . . . . . . .<br />

508 H Salix myrsinifolia Salisb. (= S. nigricans Sm.) X X X X . . . .<br />

509 Salix purpurea L. X X X X X X X .<br />

5<strong>10</strong> Salix purpurea subsp. angustior Lautenschlager X . . . . . . .<br />

511 Salix purpurea L. x S. daphnoides Vill. . . X X . . . .<br />

512 Salix purpurea subsp. gracilis (Wimm.) Buser X . . . . . . .<br />

513 Salix reticulata L. X . . . . . . .<br />

514 H Salix x rubens Schrank . . . X . X X X<br />

515 Salix triandra L. . X X . . X X X<br />

516 H Salix viminalis L. . . . . . . X .<br />

517 Salvia glutinosa L. X X X X . . . .<br />

Mauria<br />

64<br />

Forni di<br />

Sotto<br />

3 4 5 6 7 8 9 <strong>10</strong> 11<br />

Amaro<br />

Cornino<br />

Spilimbergo<br />

Casarsa<br />

Bolzano<br />

Latisana<br />

Estuary<br />

Flambro<br />

Vivaro


Investigation Areas: 1 2<br />

No. H Species a b a b a b a b a b a b a b a b a b a b a b<br />

518 H Salvia verticillata L. . . . X . . . .<br />

519 Sambucus nigra L. . X X X . . X .<br />

520 Sanguisorba minor Scop. . X X X X X X .<br />

521 Sanguisorba minor subsp. polygama (Waldst. & Kit.) . . . . . X . .<br />

522 Saponaria officinalis L. . . X X X . . .<br />

523 Satureja montana subsp. variegata (Host) P.W.Ball . . . X X . X .<br />

524 Saxifraga aizoides L. . X . . . . . .<br />

525 Saxifraga caesia L. X X . . . . . .<br />

526 Scabiosa columbaria L. X X X X . X . .<br />

527 H Scabiosa graminifolia L. . X X X X X . .<br />

528 H Scabiosa triandra L. (= Scabiosa gramuntia L.) . X . X X X X X<br />

Schoenoplectus mucronatus (L.) Palla (= Scirpus m. (L.)<br />

529 H<br />

Palla)<br />

. . . . . . X .<br />

530 Scirpus triqueter (L.) Palla . . . . . . X .<br />

Scrophularia canina L. (incl. S. juratensis<br />

531 H<br />

Schleich. ex Wydler)<br />

X X X X X X X .<br />

532 Scutellaria galericulata L. . . . . . X . .<br />

533 Sedum sexangulare L. emend. Grimm (= S. mite Gilib.) . . X X . X . .<br />

534 Selaginella helvetica (L.)Spring . X . X X . . .<br />

535 Selaginella selaginoides (L.)PB. ex Schrank & Mart. X . . . . . . .<br />

536 H Senecio erraticus Bertol. . . . . X . X .<br />

537 H Senecio inaequidens DC. . . X X X X . .<br />

538 Senecio vulgaris L. . X . . X X X .<br />

539 Seseli gouanii Koch . . X X . . X .<br />

540 H Sesleria varia Jacq. X X X X X . X .<br />

541 Setaria pumila (Poiret) Schultes . . . . . . X .<br />

542 Setaria verticillata (L.) PB. . . X X . . X .<br />

543 H Setaria viridis (L.) PB. . . X X X X X .<br />

544 Silene alba (Mill.) E.H.L.Krause . . . . . . X .<br />

545 H Silene pusilla W. & K. X X . . . . . .<br />

546 Silene vulgaris (Moench) Garcke . X X X X X . .<br />

547 Silene vulgaris subsp. angustifolia (Miller) Hajek . . . . . . . X<br />

Silene vulgaris subsp. glareosa (Jord.) Marsden-<br />

548 H<br />

Jones & Turrill<br />

X X X X . . . .<br />

549 H Silene vulgaris (Moench) Garcke subsp. vulgaris . X X X X X X .<br />

550 Sinapis arvensis L. . X . . . . . .<br />

551 Solanum dulcamara L. . X X X . . X .<br />

552 Solanum nigrum L. emend. Mill. subsp. nigrum . . X X X X X .<br />

553 Solanum nigrum subsp. schultesii (Opiz) Vessely . . . . . . . X<br />

554 Solidago canadensis L. . . . X . . . .<br />

555 Solidago gigantea var. seroti na (O.Kuntze) Cronq. . . X X X X X X<br />

556 Solidago virgaurea L. X . X X . . . .<br />

557 Sonchus arvensis L. . X X X . . X .<br />

558 Sonchus asper (L.) Hill . . X X . X X .<br />

559 Sonchus oleraceus L. . X . X . X X .<br />

560 Sorbus aria (L.) Crantz X . X . . . . .<br />

561 Sorbus aucuparia L. s.l. . X . . . . . .<br />

562 Sorbus chamaemespilus (L.) Crantz X . . . . . . .<br />

563 H Sorghum halepense (L.) Pers. . . . . X X X X<br />

564 Sparganium emersum Rehm subsp. emersum . . . X . . . .<br />

565 Spirea douglasii Hooker . . . . . . X .<br />

566 H Stachys alopecurus (L.) Bentham X . . . . . . .<br />

567 Stachys sylvatica L. . X . X . . . .<br />

568 Stellaria media (L.) Vill. . X . . . . . .<br />

569 H Stellaria nemorum L. . X X . . . X .<br />

570 H Stipa eriocaulis Borbás . . . . X . . .<br />

571 Symphytum officinale L. . . X X . . . .<br />

572<br />

Symphytum officinale subsp. bohemicum (F.W.Schmidt)<br />

Celak.<br />

. . . . . . X .<br />

573 Symphytum tuberosum subsp. nodosum (Schur) Soó . X . . . . . .<br />

574 H Tamus communis L. . . X X . X X .<br />

575 Tanacetum vulgare L. . . X X X X X X<br />

Mauria<br />

65<br />

Forni di<br />

Sotto<br />

3 4 5 6 7 8 9 <strong>10</strong> 11<br />

Amaro<br />

Cornino<br />

Spilimbergo<br />

Casarsa<br />

Bolzano<br />

Latisana<br />

Estuary<br />

Flambro<br />

Vivaro


Investigation Areas: 1 2<br />

No. H Species a b a b a b a b a b a b a b a b a b a b a b<br />

576 Taraxacum officinale L. s.l. (sect. Rud. ) . X X X X X X .<br />

577 Teucrium chamaedrys L. . . . . . . . .<br />

578 Teucrium montanum L. X X . . . . . .<br />

579 Thalictrum aquilegiifolium L. X . X X X . . .<br />

580 Thalictrum lucidum L. . . . . . . . .<br />

581 Thalictrum minus L. agg. . . X . . . . .<br />

582 Thesium divaricatum Jan ex Mert. & Koch . . . X X . . .<br />

583 Thesium linophyllon L. . X . X X . X .<br />

584 Thesium pyrenaicum Purr. . X . . . X . .<br />

585 Thesium rostratum Mert. & Koch . X X . X . . .<br />

586 Thlaspi arvense L. . . . . . X . .<br />

587 H Thymus longicaulis C.Presl . . . X . X . .<br />

Thymus praecox subsp. polytrichus (Kern. ex<br />

588 H<br />

Borb.)Ronn.<br />

. X X X X X X .<br />

589 Thymus polytrichus Kerner . . . . . X X .<br />

Thymus pulegioides L. s.l. (nach Pignatti: T. alpestris<br />

590 H<br />

Tausch)<br />

. X X X X X . .<br />

591 Tilia cordata Mill. . . . X . . . .<br />

592 Tilia platyphyllos Scop. . . X X . X . .<br />

593 Tofieldia calyculata (L.) Wahlenb. X X X . . . . .<br />

594<br />

Tolpis staticifolia (All.) Schultz-Bip. (= Hieracium<br />

staticifolium All.)<br />

. X . . . . . .<br />

595 Tortella tortuosa (L.) Limpr. . X . X . . . .<br />

596 Tragopogon pratensis L. . . X X . . . .<br />

597 Trifolium campestre Schreb. . . X X . X . .<br />

598 Trifolium medium L. . X . . . . . .<br />

599 Trifolium montanum L. X . . . . . . .<br />

600 Trifolium pratense L. . X X X X . X X<br />

601 Trifolium repens L. . . . X X X X X<br />

Trisetum argenteum (Willd.) Roem. & Schultes (T.<br />

602 H<br />

districhophyllum agg.)<br />

X X . . . . . .<br />

603 Trollius europaeus L. X . . . . . . .<br />

604 Tussilago farfara L. X X X X X X X .<br />

605 Typha latifolia L. . . X . . . X .<br />

606<br />

Ulmus glabra Huds. (= U. montana With., = U. scabra<br />

Mill.)<br />

. . X X . . . .<br />

607 Urtica dioica L. . X X X . . X .<br />

608 Vaccinium vitis-idaea L. X . . . . . . .<br />

609 Valeriana montana L. X . . . . . X .<br />

6<strong>10</strong> Valeriana officinalis agg. . . X . . . . .<br />

611 Valeriana saxatilis L. X X . . . . . .<br />

612 Verbascum densiflorum Bertol. . . . . . X . .<br />

613 Verbascum thapsus L. (evtl. V. thapsiforme Schrader?) . . X . . . . .<br />

614 Verbena officinalis L. . . . . . X X .<br />

615 Veronica anagallis-aquatica L. . X X X X X X .<br />

616 H Veronica beccabunga L. . X X X . X X .<br />

617 Veronica persica Poir. . X . X . . . .<br />

618 Veronica urticifolia Jacq. X X . . . . . .<br />

619 Viburnum lantana L. X X X X . . . .<br />

620 Viburnum opulus L. . X X X . . . .<br />

621 Vicia cracca L. X X X X X . . .<br />

622 Vincetoxicum hirundinaria L.W.Medicus s.l. X X X . . . . .<br />

623 Vincetoxicum hirundinaria L.W.Medicus subsp. . . . X . . . .<br />

624 Viola biflora L. X . . . . . . .<br />

625 Viola hirta L. X X . . . . . .<br />

626 Viola riviniana Rchb. . X X X . . . .<br />

627 Vitis vinifera L. . . X X . . X .<br />

628 H Xanthium saccharatum Wallr. (= X. italicum . . X X X X X .<br />

629 H Xanthium strumarium L. . . X X X . X .<br />

630 . . . . . . . .<br />

631 . . . . . . . .<br />

632 . . . . . . . .<br />

Mauria<br />

66<br />

Forni di<br />

Sotto<br />

3 4 5 6 7 8 9 <strong>10</strong> 11<br />

Amaro<br />

Cornino<br />

Spilimbergo<br />

Casarsa<br />

Bolzano<br />

Latisana<br />

Estuary<br />

Flambro<br />

Vivaro


8 LITERATURE<br />

8.1 GENERAL LITERATURE:<br />

ADAC Verlag GmbH (1995): Das große ADAC Alpenbuch. München und Mairs<br />

Geographischer Verlag, Ostfildern<br />

Adler, W., Oswald, K. & Fischer, R. (1994): Exkursionsflora von Österreich. Verlag Eugen<br />

Ulmer, Stuttgart und Wien<br />

Barndt, D., Platen, R. & Wachmann, E. (1995): Laufkäfer: Beobachtung und Lebensweise.<br />

Naturbuchverlag Augsburg<br />

Bellmann, H, (1993): Heuschrecken; Beobachten<strong>–</strong>Bestimmen. Naturbuchverlag Augsburg<br />

Bunalski, M. (1999): Die Blatthornkäfer Mitteleuropas, Coleoptera, Scarabaeoidea;<br />

Bestimmung <strong>–</strong> Verbreitung <strong>–</strong> Ökologie. Slamka, Bratislava<br />

European Commission DG Environment (2007): Interpretation Manual of European<br />

Habitats (EUR 27)<br />

Fontana, P., Buzzetti, F. M., Cogo, A. & Baudewijn, O. (2002): Guida al Riconoscimento e<br />

allo studio di Cavallette, Grilli, Mantidi e Insetti affini del Veneto. Blattaria, Mantodea,<br />

Isoptera, Orthoptera, Phasmatodea, Dermaptera, Embiidina. Museo Naturalistico<br />

Archeologico di Vicenza Ed., Vicenza<br />

Geigenmüller, K. & Trautner, J., (1987): Sandlaufkäfer, Laufkäfer: Illustrierter Schlüssel zu<br />

den Cicindeliden und Carabiden Europas. J. Margraf, Aichtal<br />

Huemer, P. (1996): Lepidopteren im Bereich der dealpinen Flüsse Medusa und<br />

Tagliamento (Friuli-Venezia Giulia, Norditalien). Gortania Atti del Museo Friulano di<br />

Storia Naturale 18: 201-214<br />

Jäger, J.E. & Werner, K. (eds.) (2005) Rothmaler <strong>–</strong> Exkursionsflora von Deutschland Band<br />

2 & 3 <strong>–</strong> Gefäßpflanzen Grund- & Atlasband. Elsevier/Spektrum Akademischer<br />

Verlag, München<br />

Kahlen, M. (2002): Die Käfer der Ufer und Auen des Tagliamento (Erster Beitrag: Eigene<br />

Sammelergebnisse). Gortania Atti del Museo Friulano di Storia Naturale 24: 147-202<br />

Kollmann, J., Viele, M., Edwards, P.J., Trockner, K. & Ward J.V. (1999): Interactions<br />

between vegetation development and island formation in the <strong>Alpine</strong> river<br />

Tagliamento. Applied Vegetation Science 2: 25-36<br />

Kuhn, K. (1995): Beobachtungen zu einigen Tiergruppen am Tagliamento. Jahrbuch des<br />

Vereins zum Schutz der Bergwelt 60: 71-86<br />

Kuhn, K.(2005): Die Kiesbänke des Tagliamento (Friaul, Italien) <strong>–</strong> Ein Lebensraum für<br />

Spezialisten im Tierreich. Jahrbuch des Vereins zum Schutz der Bergwelt 70: 37-44<br />

Kunth, Travelmag Reiseatlas (2005/2006): Reiseatlas Alpen, Provence, Cote d´Azur,<br />

Norditalien, Europa. GeoGraphic Puplisher GmbH & Co.Kg, München<br />

Lautenschlager-Fleury, D. & E. (1994): Die Weiden von Mittel- und Nordeuropa;<br />

Bestimmungsschlüssel und Artenbeschreibung für die Gattung Salix L. Birkenhäuser<br />

Verlag, Basel, Bosten und Berlin<br />

Lippert, W., Müller, N., Rossel, S., Schauer, T. & Vetter, G. ( 1995 ): Tagliamento-<br />

Flußmorphologie und Auenvegetation der größten Wildflußlandschaft in den<br />

Alpen. Jahrbuch des Vereins zum Schutz der Bergwelt 60: 11-70<br />

67


Mertz, P. (2000): Pflanzengesellschaften Mitteleuropas und der Alpen; Erkennen,<br />

Bestimmen, Bewerten, Ein Handbuch für die vegetationskundliche Praxis. ecomed<br />

Verlagsgesellschaft, AG & Co. KG<br />

Müller, N. & von Heßberg, A. (2005): Alpenworkshop und <strong>International</strong>e Flusskonferenz<br />

Tagliamento 2005 <strong>–</strong> Ergebnisse. Selbstverlag Fachbereich Landschaftsarchitektur<br />

FHE, Erfurt<br />

Müller, N. (1991a): Veränderungen alpiner Wildflußlandschaften in Mitteleuropa unter dem<br />

Einfluß des Menschen. Augsburger Ökologische Schriften 2: 9-30<br />

Müller, N. (1991b): Auenvegetation des Lech bei Augsburg und ihre Veränderungen<br />

infolge von Flußbaumaßnahmen. Augsburger Ökologische Schriften 2: 79-<strong>10</strong>8<br />

Müller, N. (1995): Wandel von Flora und Vegetation nordalpiner Wildflußlandschaften<br />

unter dem Einfluß des Menschen. Ber. ANL 19: 125-187<br />

Müller, N. (2004): Skriptum: Vegetationsökologie. Fachbereich Landschaftsarchitektur<br />

FHE, Erfurt n.p.<br />

Müller, N. (2005): Die herausragende Stellung des Tagliamento (Friaul, Italien) im<br />

Europäischen Schutzgebietssystem NATURA 2000. Jahrbuch des Vereins zum<br />

Schutz der Bergwelt 70: 19-35<br />

Müller, N., Besch, T., Grosser, N. & Müller, O. (2006): Materialien zum <strong>International</strong>en<br />

Alpen <strong>Workshop</strong> Tagliamento 2006. Selbstverlag Fachbereich<br />

Landschaftsarchitektur FHE, Erfurt<br />

Müller, N., Dalhof, B., Häcker, B.& Vetter, G. (1992): Auswirkungen unterschiedlicher<br />

Flußbaumaßnahmen auf Flußmorphologie und Auenvegetation des Lech - eine<br />

Bilanz nach <strong>10</strong>0 Jahren Wasserbau. Ber. ANL 16: 181-214<br />

Pfadenhauer, J. (1993): Vegetationsökologie <strong>–</strong> ein Skriptum. IHW, Eching<br />

Pfeuffer, E. (2003): Zur Heuschreckenfauna des Tagliamento (Norditalien). Articulata<br />

18(2): 215-225<br />

Poldini, L. (1991): Atlante corologico delle piante vascolari; Nel Friuli-Venezia Giulia <strong>–</strong><br />

Inventario floristico regionale. Udine<br />

Poldini, L., Oriolo, G., Vidali, M. (2001): Vascular Flora of Friuli-Venezia Giulia. An<br />

annotated Catalogue and synonimic Index. Studia Geobotanica 21(1): 1-227<br />

Reich, M. (1991): Grasshoppers (Orthoptera, Saltatoria) on alpine and dealpine riverbanks<br />

and their use as indicators for natural floodplain dynamics. Regulated rivers:<br />

Research and Management, Vol. 6, 333-339<br />

Reisigl, H. & Keller, R (1987): Alpenpflanzen im Lebensraum; <strong>Alpine</strong> Rasen Schutt- und<br />

Felsvegetation. Gustav Fischer Verlag, Stuttgart<br />

Stresemann, E. (2000): Exkursionsfauna von Deutschland; Wirbellose: Insekten. Spektrum<br />

Akademischer Verlag Heidelberg<br />

Tockner, K., Surian, N. & Toniutti, N. (2005): Geomorphologie, Ökologie und nachhaltiges<br />

Management einer Wildflusslandschaft am Beispiel des Friume Tagliamente (Friaul,<br />

Italien) <strong>–</strong> ein Modellökosystem für den Alpenraum. Jahrbuch des Verein zum Schutz<br />

der Bergwelt 70: 3-17<br />

Westermann Schulbuchverlag GmbH (1991): Dierke Weltatlas. Westermann Druck,<br />

Braunschweig<br />

68


8.2 WORKING LITERATURE AND MAPS<br />

Tab. 11: Working literature and maps inclusive number of copies<br />

Nr. Author(s) Year Title Publisher Owner Quantity<br />

1<br />

Adler, W., Oswald, K. & Fischer,<br />

R.<br />

1994 Exkursionsflora von Österreich<br />

Verlag Eugen Ulmer,<br />

Stuttgart und Wien<br />

WL 1124 A237 4<br />

2<br />

Aeschimann, D., Lauber, K.,<br />

Moser, D. M., Theurillant, J.-P.<br />

2004 Flora alpina, Band 1<br />

Verlag Paul Haupt, Bern,<br />

WL 1125 A253-1<br />

Stuttgart, Wien<br />

1<br />

3<br />

Aeschimann, D., Lauber, K.,<br />

Moser, D. M., Theurillant, J.-P.<br />

2004 Flora alpina, Band 2<br />

Verlag Paul Haupt, Bern,<br />

WL 1125 A253-2<br />

Stuttgart, Wien<br />

1<br />

4<br />

Aeschimann, D., Lauber, K.,<br />

Moser, D. M., Theurillant, J.-P.<br />

2004 Flora alpina, Band 3<br />

Berichte dea Instituts für Landschafts- und<br />

Verlag Paul Haupt, Bern,<br />

WL 1125 A253-3<br />

Stuttgart, Wien<br />

1<br />

5<br />

Arndt, U., Böcker, A., Kohler, A.,<br />

Konold, W., Müller, W.A.<br />

1996<br />

Pflanzenökologie der Universität<br />

Hohenheim- Flüsse und<br />

Flusslandschaften in Friaul-Veneetien,<br />

Slowenien und Kroatien<br />

Norbert Müller 1<br />

6 Comitato Studi Tagliamento. 2005<br />

Tagliamento due sponde sul fiume- Guida<br />

Comitato studi<br />

storico tecnica di un tratto del medio<br />

Norbert Müller<br />

Tagliamento Spilimbergo<br />

corso<br />

Invasive Neophyten am Tagliamento<br />

1<br />

7 Geobotanisches Institut Zürich.<br />

2005/0<br />

6<br />

(Italien)- Bestandsaufnahme,<br />

Invasionsprozesse auf Inseln, Ökologie<br />

von Amorpha fructicosa<br />

Norbert Müller 1<br />

8 Heßberg, A., Klotzenburg, M. 2009<br />

Tagliamento Exkursion- Universität<br />

Tübingen und Universität Hohenheim<br />

Norbert Müller 1<br />

9 Klapp, E., Opitz von Boberfeld,W. 1990 Taschenbuch der Gräser. 12. Auflage<br />

Verlag Paul Parey,<br />

Berlin und Hamburg<br />

WL9680 K63 (12)<br />

+7<br />

1<br />

<strong>10</strong> Lauber. K,. Wagner, G. 2001 Flora helvetica 3. Auflage<br />

Verlag Paul Haupt, Bern,<br />

WL 1126 L366 (3)<br />

Stuttgart, Wien<br />

1<br />

11<br />

Lippert, W., Müller, N., Rossel,<br />

S., Schauer, T., Vetter, G.<br />

1995<br />

Der Tagliamento- Flußmorphologie und<br />

Auenvegetation der großen<br />

Wildflusslandschaften in den Alpen<br />

Norbert Müller 2<br />

12 Kuhn,K. 1995<br />

Beobachtungen zu einigen Tiergruppen<br />

am Tagliamento<br />

Norbert Müller 1<br />

13 Kretschmer, W. 1995<br />

Hydrobiologische Untersuchungen am<br />

Tagliamento (Friaul, Italien)<br />

Norbert Müller 1<br />

14 Mion, B. 2005<br />

Il Tagliamento- Immagin eappunti sul "rei<br />

del fiumi alpini"<br />

Verlag<br />

ediRISMA,Roveredo in<br />

Piano<br />

Eigenverlag FHE -<br />

RO 45915 M 669 1<br />

15 Müller, N. 2004 Alpen <strong>Workshop</strong> Tagliamento 2004 Fachbereich<br />

Landschaftsarchitektur<br />

Norbert Müller 1<br />

16<br />

Müller, N., Bangert, U., Drecher,<br />

A., Kuhn, K.<br />

2004<br />

Alpen <strong>Workshop</strong> Tagliamento 2004-<br />

Ergebnisse<br />

Eigenverlag FHE -<br />

Fachbereich<br />

Landschaftsarchitektur<br />

Norbert Müller 1<br />

Alpen <strong>Workshop</strong> & Inernationale Eigenverlag FHE -<br />

17 Müller, N., Heßberg, A. 2005 Flusskonferenz Tagliamento 2005 - Fachbereich<br />

Norbert Müller 1<br />

Ergebnisse<br />

Landschaftsarchitektur<br />

18<br />

Müller, N., Heßberg, A.,<br />

Radenberg, E.<br />

2005<br />

Materialien zum Alpen <strong>Workshop</strong> & zur<br />

<strong>International</strong>en Flusskonferenz<br />

Tagliamento 2005<br />

Eigenverlag FHE -<br />

Fachbereich<br />

Landschaftsarchitektur<br />

Norbert Müller 1<br />

19<br />

Müller, N. , Verein zum Schutz<br />

der Bergwelt e.V.<br />

2005<br />

Rettet den Tagliamento Friaul/Italien-<br />

König der Alpenflüsse<br />

Norbert Müller 2<br />

20 Poldini, L. 1991<br />

Atlante corologicol delle Piante vascolari<br />

nel Friuli-Venezia Giulia<br />

Inventario floristico<br />

regionale. Udine<br />

WL 1136 P762 1<br />

21 Schiechtl. H.M. 1992<br />

Weiden in der Praxis- Die weide<br />

Mitteleuropas, ihre Verwendung und ihre<br />

Bestimmung<br />

Verlag Patzer, Berling-<br />

Hannover<br />

WL 8865 S332 +4 4<br />

22 Rossel, S. 1993<br />

Tagliamento / Friaul (Nordost-Italien) -<br />

Ergebnisse der Exkursion<br />

Norbert Müller 1<br />

23 Tobacco ?<br />

Carta topographica per escursionisti<br />

M1:25.000. Foglio 02,013, 020,020, 027<br />

Norbert Müller je 1<br />

24 Tobacco 2005<br />

Carta stradale della Regione autonoma<br />

Friuli-Venezia Giuglia M 1:<strong>10</strong>0.000<br />

Norbert Müller 1<br />

69


9 LEGEND TO PICTURES ON FRONT & BACK COVER<br />

Front cover outside:<br />

1. 1991 Cornino<br />

Front cover inside:<br />

2. 2005 demonstration<br />

for preservation of<br />

Tagliamento at<br />

Gemona<br />

4. 2007 Mt. Ragogna, upstream view<br />

70<br />

3. 2005 1 st River<br />

conference at<br />

St. Pietro<br />

1. 2006 Gemona 2. 2009 Forni di Sotto 3. 2007 Cornino<br />

Back cover inside:<br />

1. 1991 Forni di Sotto<br />

3. 1992 Bibione<br />

5. 2005 Passo della Mauria, a<br />

presentation by Andreas von<br />

Heßberg<br />

7. 2005 final ceremony at Gemona<br />

9. 2006 Spilimbergo<br />

Back cover outside:<br />

1. 2006 Forni di Sotto<br />

3. 2007 Mt. Ragogna<br />

5. 2009 Passo della Mauria<br />

7. 2009 Bolzano<br />

9. 20<strong>10</strong> Vivaro<br />

2. 1991 Mt. Ragogna<br />

4. 2004 Cornino<br />

6. 2005 Bolzano, a presentation by<br />

Manfred Kahlen<br />

8. 2005 final ceremony at Gemona<br />

<strong>10</strong>. 2006 Amaro, Mantis religiosa L.<br />

2. 2007 Casarsa<br />

4. 2007 at Hotel Ai Glicini, Cornino<br />

6. 2009 Cornino<br />

8. 20<strong>10</strong> Flambro<br />

<strong>10</strong>. 20<strong>10</strong> Tauern/Alps<br />

All Photographs belong to their respective owners & Tagliamento <strong>Workshop</strong> 1991-2011.

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