18 Bulletin <strong>of</strong> the <strong>California</strong> Znsecf Survey Map 10. Locality records in <strong>California</strong> for Ctenophora (Phoroctenia) vittata angustipennis Loew (circles). CIS). Berkeley, Strawberry Canyon, XI-16-63, larvae in rotting logs <strong>of</strong> red alder (Alnus oregona) (A. G. Raske and R. Rollins, CIS). Berkeley, Woolsey Canyon, V-1148 (J. W. MacSwain, CIS). Mills College, 111-9/12-26 (E. Craig, CAS). Oakland, from Querws agrifoliaD 111-27-52 (R 0. Schuster, CIS). LAKE Co.: Anderson Springs, III-30-61 (J. S. Buckett, UCD) . MARIN CO. : Kentfidd, ?EdyptudJD 111-18-23 (CIS). Kent Lake, 11-27-60 (C. A. Toschi, CIS). Lagunitas Creek, IV-15-1876 (Osten Sacken, MCZ). Mill Valley, V-14-41 (D. Q. Cavagnaro, UCD). Mill Valley, Cascade Creek, IV-1- 50, V-11-52 (H. B. Leech, CAS). Mill Valley, Falls Creek, emerged from dead Lithocarpus densiflorus, IV-25-57 (H. B. Leech, CAS). Samuel P. Taylor State Park, IV453 (E. S. Ross, CAS). San Rafael, II-1935 (C. D. Duncan, SJS). MONTEREY CO.: Santa Lucia Mountamq Hastings Reservation, IV-19-38 (J. Linsdale, CIS). SAN BERNARDINO CO.: Arrowhead Lake, VI-3-28 (E. C. Van Dyke, CAS). SAN MATEO CO.: 2 mi. NE <strong>of</strong> La Honda, larvae and pupae in well-rotted alder, 11-29-64 (J. T. Doyen, CIS). SANTA CLARA Co.: Monta Vista, near post <strong>of</strong>fice, north side <strong>of</strong> Stevens Creek, in Santa Clara valley, reared from oak log, 111-2944 (T. L. Erwin, SJS). Palo Alto, IV-1915 (R. W. Doane, CAS). SANTA CRUZ CO.: Santa Cru~, 11-23-29 (E. P. Van DUZ- CAS); 111-30-61 (R. M. Brown. CAS.) Waddell Creek, 111-8-30, IV-5/10-30, V-130 (F. Bianchi, CAS). SHASU Co.: Shingletown, VI-2-41 (C. W. Anderson, CIS). SONOMA Co.: Healdsburg, Mill Creek, IV-1/15-50 (R. E. Leech, CAS). Trinity, IV4-37 (N. W. Frazier, CIS). Genus Nephrotoma Meigen Pachyrrhina Macquart, 1834, Hist Nat Ins., Dipt 1:88 (variant spellings, Pa~hphi~~ Pachyrhyna). Nephr<strong>of</strong>oma is one <strong>of</strong> the largest genera in the Tipuline crane flies, being best developed in the Holarctic, Oriental, and Ethiopian regions, with about 40 species recorded from the Nearctic region and in the 1965 Catalog <strong>of</strong> the Diptera. Of these, it appears that a single species occurs in <strong>California</strong>, the prior name being wulpiana (Bergroth). This was described from a single female specimen taken in Siskiyou County by Behrens and has not been recognized in recent collections. <strong>The</strong> later described occidentalis (Doane) is abundant in the state and while differing in minor details <strong>of</strong> coloration from Bergroth?s description <strong>of</strong> wulpiana, appears certainly to represent the same species and therefore is placed in the synonymy. In case future investigations should indicate that two distinct species are involved, all records here listed should be referred to occidentalis with the single exception <strong>of</strong> Bergroth?s type specimen <strong>of</strong> wulpiana (Siskiyou county, below). A second very distinct species, Nephrotorna alfissima (Osten Sacken), is western, having been taken in Nevada. It is almost entirely black or, in the color form, eryfhrophrys (Williston) with the abdomen chiefly orange, the hypopygium black. 1 Pales Meigen, 1800, Nouv. Class. Mouch., p- 14 (nom. nud.). Nephmtoma Meigen, 1803, Illiger?s Mag. 3:262. Map 11. Locality records in <strong>California</strong> for Nephrotoma wulpiana (Bergroth) (circles).
Nephrotoma wulpiana (Bergroth) (Figs. 9, 50; Map 11) Pachyrrhina wulpiana Bergroth, 1888, Wien. Ent. Ztg. 7: 200. Holotype 0; Siskiyou County, Calif. (location <strong>of</strong> type not known). Synonym.-Pachyrhina occidentalis Doane, 1908, Ent News 19: 177-178. Syntypes 8 9; <strong>California</strong> and other western states (Stanford University, now CAS). Geographic range-British Columbia; Arizona, <strong>California</strong>, Montana, Nevada, Oregon, Texas, Utah, Washington. <strong>California</strong> records.-ALAMEDA CO.: Berkeley, university <strong>of</strong> <strong>California</strong> campus, IV-6-50 (J. W. MacSwain, CIS); V-5,16- 58 (D. J. Burdick, CIS); V-16-58 (W. A. Nisbet, CIS). Oakland, IV-20-41 (D. A. DeCass, CIS); V-10-50 (L. Bowers, CIS). Oakland Hills, XI-1938 (CAS). COLUW Co.: Coluw, VIXX-15-55 (R. 0. Schuster, CIS). CONTRA COSTA CO.: Moraga, VI-13-65, X-14-65 (D. G. Denning, CPA). Walnut Creek, (foot <strong>of</strong> Shell Ridge), 111-10-63, IX-24/26-63 (J. A. Powell, CIS). FRESNO CO.: Lost Lake, 1 mi. W <strong>of</strong> Friant, VIX-1-56 (R. 0. Schuster, CIS). HUMBOW Co.: Big Lagoon, VIII-12-63 (J. S. Buckett, UCD). INYO m.: Bishop, 4500 feet, IV-16-64 (J. D. Birchim, CPA). Camp Manaanar, 3800 feet, VII-1/6-?5 (J. M. Mayeda, CPA). Independence, N fork <strong>of</strong> Oak Creek above fish hatchery, 5000 feet, VI-27-63 (C. P. Alexander, CPA) ; 4000 feet, V-12-64 (J. D. Birchim, CPA). Lone Pine, 3800 feet, V-15-64 (J. D. Birchim, CPA). Oak Creek fish hatchery, 3950 feet, V-16-63 (C. P. Alexander, CPA). KERN CO.: Bakersfield, in house, IV-1961 (R. E. Bellamy, CPA). Greenhorn Mountains, Tributary '93," Cedar Creek, V-15-61 (R. E. Bellamy, 5020, CPA). Kernville, along Kern River, V-13-63 (C. P. Alexander, CPA). Los ANGELES CO.: Lancaster, in alfalfa field, VII-28-58 (E. I. Schlinger, UCR). MARIN CO.: Mill Valley, at light, VIII-19-50 (H. B. Leech, CAS). MONO Co.: Coleville, V-27-39 (T. H. G. Aitken, CPA). MON'I'EREY CO.: Kirk Creek, WI-10-62 (E. I. Schlinger, UCR). ORANGE Co.: Hot Springs Canyon, between Capistrano and Elsinore, IV-1-63 (C. P. Alexander, CPA). RIVERSIDE CO.: Cleveland National Forest, Santa Ana Mountains, San Juan Canyon, Upper San Juan Forest Camp, 1745 feet, V-8-63 (C. P. Alexander, CPA). Riverside, IV- 13-26 (P. H. Timberlake, CPA). White Water, 111-14-34 (P. H. Timberlake, CPA). SAN BERNARDINO CO.: San Bernardino Mountains, Forest Home, 5100 feet, V-9-63 (C. P. Alexander, CPA); Mill Creek Camp, Thurman Flats picnic area, 3400 feet, V-10/26-63 (C. P. Alexander, CPA); Seven Oaks, along Santa Ana River, 5000 feet, V-12-63 (C. P. Alexander, CPA). SAN DIEGO CO.: Cleveland National Forest, outlet <strong>of</strong> El Capitan Lake, about 7 mi. E <strong>of</strong> Lakeside, 700 feet, IV-12-63 (C. P. Alexander, CPA). Del Mar, IV-18-63 (C. P. Alexander, CPA). La Jolla, VI-22-63 (R. L. Langston, CIS); VII-17,19-63, on Eriogonum fasciculafum, (P. D. Hurd, CIS); VI-17-63 (I. H. Pogojeff, UCD). Palomar Mountain, Doane Creek, 4700 feet, VIE745 (J. A. Comstock, CPA). SAN FRANCISCO CO.: South San Francisco, 111-21-50 (R. S. Beal, CIS). SAN LUIS OBISPO CO.: Atascadero, V-3- 38 (G. S. Mansfield, SJS). SAN MATEO CO.: Menlo Park, IV-18-62 (P. J. Gagen, SJS). Redwood City, 1952 (P. H. Arnaud, CPA). SANTA CLARA CO.: Palo Alto (Doane); part <strong>of</strong> type material <strong>of</strong> occidentalis. San Jose, IV-13-58 (D. Burdick, CIS). SHASTA CO.: Castle Crags State Park, along Sacramento River, 2000 feet, VII-28-50 (C. P. Alexander, Alexander: <strong>The</strong> <strong>Crane</strong> <strong>Flies</strong> <strong>of</strong> <strong>California</strong> 19 CPA). SISKIYOU Co.: Without more exact data (J. Behrens); type <strong>of</strong> wulpiana, location <strong>of</strong> type specimen unknown. TULARE CO.: Camp Nelson, Nelson Creek, 5000 feet, V-29-63 (C. P. Alexander, CPA). Fairview, along Kern River, V-13-63 (C. P. Alexander, CPA). Three Rivers, along Kaweah River, V- 30-63 (C. P. Alexander, CPA). Tule River at east end <strong>of</strong> Success Lake, V-19-63 (C. P. Alexander, CPA). Genus Tipula Linnaeus Tipda Linnaeus, 1758, Syst. Nat Ed. 10:585. Tipula is one <strong>of</strong> the major groups <strong>of</strong> insects, including nearly one-sixth <strong>of</strong> all known crane flies and, in <strong>California</strong>, nearly 40 percent <strong>of</strong> the species. <strong>The</strong> genus has been divided into slightly more than 40 subgeneric groups, <strong>of</strong> which 20 are found in the state. <strong>The</strong> most recent listing <strong>of</strong> the Nearctic subgenera, together with a general discussion <strong>of</strong> the structure <strong>of</strong> the male hypopygium, is included in a paper by the author (Alexander, 1965). In the accompanying key to the subgenera, the presence or absence <strong>of</strong> strong setae on the squama or axillary lobe <strong>of</strong> the wing is utilized. It has been necessary to use characters <strong>of</strong> the male hypopygium in defining several <strong>of</strong> the subgeneric groups. <strong>The</strong>se and their included species have been arranged alphabetically. However, in the large and involved subgenus Lunatipula Edwards, where 20 species groups are recognized, the figures illustrating the genitalia are arranged by groups in order to facilitate comparison <strong>of</strong> related species. <strong>The</strong> following subgenera are represented in <strong>California</strong> by single species: Angarotipula, Arctotipula, Bellardina, Labiotipula, Lindnerina, Odonatisca, Platytipula, Savtshenkia, and Tipulodindes. <strong>The</strong> subgenera Platytipula, Tipula, and Vestiplex are regional. KEY TO THE SUBGENERA OF TIPULA Antennal flagellar segments without verticils (fig. 10); (squama naked) . ANGAROTIPULA Savchenko illustris Doane Antennal flagellar segments with conspicuous verticils (fig. 11) . *... . . . . . . . . . 2 Outer cells <strong>of</strong> wing with macrotrichia 3 No macrotrichia in wing cells . . . . . . . . 5 Coloration <strong>of</strong> thorax polished orange or yellow, in cases with contrasting black or darkened stripes (separate key, p. 27) . HESPEROTIPULA Alexander in part Coloration <strong>of</strong> thorax opaque yellow, brown, or gray, darkened stripes opaque . . . . , . . . . 4 Squama <strong>of</strong> wing naked; male hypopygium with posterior border <strong>of</strong> tergite with a U-shaped or V- shaped notch, the lobes with microscopic blackened setoid spicules (lacking in gertschi) ; aedeagus with penis tube elongate and unusually stout (figs. 210-225) (separate key, p. 56) . . . . . . TRICHOTIPULA Alexander in part Squama <strong>of</strong> wing with setae; male hypopygium with posterior border <strong>of</strong> tergite broadly and very shallow-
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