The Crane Flies of California - Essig Museum of Entomology ...

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32 Bulletin of the California Znsect Survey California records.-HUMBOLDT Co.: Prairie Creek State Park; VII-11-53 (C. P. Alexander, CPA). MARIN CO.: Fairfax, IV-13-19 (CPA). SANTA CRUZ CO.: Smta CrUZ, VI-1,2-19 (CPA). Tipula (Hesperotipula) supplicata Alexander (Fig. 115) Tipula (Lunatipula) aupplicata Alexander, 1944, Canad Ent 76: 170-171. Holotype 8; San Jacinto River Canyon, Riverside County, Calif. (CPA). Geographic raztge.-Known only from California. California recOrds.--KBm CO.: 7 mi HIO Point, V-9-61 (R. E. Bellamy, 5017, CPA). RMCRSILJE Co.: San JaCinto Mountains, Idyllwild, V-14-39 (T. H. G. Aitken, -A); type. Tipula (Hesperotipula) sweetae Alexander (Fig. 116) Tipula sweetae Alexander, 1930, Brooklyn Ent Soc. Bd. 25:278-279. Holotype 8 ; Claremont, Los Angeles County, Cali. (CPA). Geographic range.-Known only from California. California recOrds.--loS ANGELES Co.: Claremont (H. E. Sweet, CPA); type. RIVERSIDE Co.: Riverside, 11-12-35 (A. L. Melander, CPA); 11-19-35. III-2/15-35, IV-1/1435 (P- H. Timberlake, CPA). SAN D ~oo Co.: Jacumba, IV-11-53 (J. A. Powell, CIS). Subgenus Labiotipula Alexander Tipula (Labiotipula) Alexander, 1965, Pacific Insecta 7:343, fig. 13. The subgenus was proposed for species of the socalled macrolabis group, previously assigned to Lunatipula. The type of the subgenus, T. (L.) macrolabis Loew, is very widespread in northern North America, extending to northeastern Asia, and may possibly be found in the state. Tipula (Labiotipula) leechi Alexander (Fig. 117; Map 22) Tipula leechi Alexander, 1938, Brooklyn En+ Soc. Bul. 33:71- 72. Holotype 8 ; Vancouver, British Columbia (CPA). Geographic range.-British Columbm; California. California records.-EL DORAW Co.: Snowliie Camp, VI- 24-48 (P. D. Hurd, CIS). PLUMAS Co.: Meadow Valley, 3500-4000 feet, VI-4-24 (E. C. van Dyke, CAS). SHASTA Co.: Burney, V-3039 (M. A. Cazier, CPA). Subgenus Lindnerina Mannheims Tipula (Lindneria) Mannheims, 1963, Stiittg. Beitr. Naturk. 102:22 (Preocc. by Lindneria Kroeber, 1929, Diptera). Tipula (Lindnerina) Mannheims, 1965, Lounais-Hbeen Luonto, 20:3. A very small group of northern Holarctic Tipula, the best known Nearctic species being the widespread T. (L.) serta Loew, occurring as far west as Alberta. A single High Sierran species occurs in California. m u ”, u 111 1 . u --. .I . I* .*+I Map 22. Locality records in California for Tipula (Labiotipula) leechi Alexander (circles), T. (Lindnerina) shieldsi Alexander (triangles), and T. (Lunatipula) amtipleura Doane (squares). Tipula (Lindnerina) shieldsi Alexander (Fig. 118; Map 22) . Tipula (findneria) shieldsi Alexander, 1965, Pacific Insects 7 : 366367. Holotype 8; Mount Dana, Mono County, Calif. (CPA). Geographic range.-Knm only from California. California records.-mo CO.: Southeast, above “L” Lake, near French Canyon, 11,600 feet, VIII-14-63 (0. Shields, CPA). South, above Golden Trout Lake, Glacier Divide, 11,500 feet, WI-12-63 (0. Shields, CPA); paratypes. FRBS”YO Cos: Piute Pass, Humphreys Basin, 11,400 feet, VIII-11-63 (0. Shields, CPA); paratypes. INYO Co.: Northwest, Mono Pass, 12,000 feet, VIII-8-61 (C. D. MacNeill, D. C. Rentz, M. R. Lundgren, CAS); paratypes. MONO Co.: North slope of Mount Dana, 11,000-11,300 feet, VIII-l-64 (0. Shields, CPA) ; holotype 8, allotype 0. Subgenus Lunetipula Edwards Tipula (Lunatipula) Edwards, 1931, Ann. and Mag. Nat Hist ser. 10, 8:81-82. Lunatipula is the largest subgenus of Tipula in the state and is one of the most difficult taxonomically. For convenience in handling the numerous species, these have been placed in 20 groups and have been arranged in the smaller units in the keys and on the
Alexander: The Crane Flies of California 33 plates showing the hypopygial structure. However, in the final listing of the species, all in the subgenus have been arranged alphabetically, as has been done with other genera and subgenera in this paper. As was indicated in the Introduction, most of the figures in the genus Tipula were drawn by the very capable young artist, Mr. Steven L. Jensen, of Salt Lake City. Relatively few were made by the author, these being chiefly of species that became available late in the preparation of the paper. 1 2 3 KEY TO THE SPECIES OF LUNATIPULA Male hypopygium with posterior border of eighth sternite produced into lateral lobes that bear a fleshy appendage, without setae (figs. 156, 159, 161) or with one or more fasciculate setae compris- ed af few ta semi bristlm fused into a unit with their filamentous tips free (sternata, fig. 160), M with one or more of the setae noticeably enlarged . Male hypopygium with none of the setae of the eighth sternite markedly enlarged or fasciculate (fie. 119, 120, 125-129, 137-155) . . . . . 30 Eighth sternite with lateral appendage glabrous, without setae (&+ 156-159, 161) or stemata (6g. 160) with several setae fasciculated to appear as a unit, with their tips free . . . . . . . . 3 I terdata group Eighth sternite with lateral appendage or lobe with distinct separate setae (figs. 121-124, 162-168, 170, 171, 174-179) . . . . . . . . . . 9 Ninth tergite produced into conspicuous lateral horns (figs. 156, 160, 161) . . . . . . . . . . 4 Ninth tergite with lobes obtuse, only slightly produced (fie. 157-159) . . . . . . . . . . 6 4 Inner dististyle with the outer basal lobe a long flattened blade, the tip obtuse, nearly as long as the body of style (fig. 156) . kirkwoodiana Alexander Inner dististyle with the outer basal lobe relatively 5 6 small and inconspicuous (figs. 160, 161) . . . 5 Lateral tergal horns slightly decurved outwardly, their tips obtuse; median tergal area extensive, the lateral end at base of horn with a blackened knob; eighth sternite with lateral lobes broad, fleshy, without setae (fig. 161) . . . . . tergafa Doane Lateral tergal horns nearly straight, tips acute; median tergal area with two intermediate pale blades; eighth sternite with lateral lobes divided, the inner arm broad with a compact fasciculate mass of setae that appears as a unit with free tips (fig. 160) . . . . . . . . sternata Doane Lateral arms of eighth sternite almost equally bilobed; lateral tergal horns narrow, median area with two projections; inner dististyle with outer basal lobe simple, not spinous; phallosome with the decurved spine simple . . . . . . . . . . . 7 Lateral arms of eighth sternite extended into a slender fingerlike lobe; lateral tergal horns broad, their apices oblique, median area not projecting; inner dististyle with outer basal lobe produced into spinous points; phallosome with the decurved spines forked at tips (figs. 157-159) . , . . . . . 8 2 7 Lateral tergal horns broader, tips obliquely truncate; median projections low and inconspicuous; inner dististyle with posterior crest not produced backward (Arizona Report, p. 500, fig. 4, as buenoi) . 8 Aavocada mane Lateral tergal horns appearing as narrow blackened points; median projections blackened, conspicuous, tips obtuse; inner dististyle with posterior crest produced backward as a flattened blade . . . . mohavensi-s Alexander (Extralimital:Arizona) Lateral tergal horns more produced; inner dististyle with the outer basal lobe modemtely large, produced into two or three strong spines; phallosome with the decurved apophyses slender, terminating in two short divergent spinous points (figs- 157, 158) . . . . . . . . . . mesotergata Alexander Lateral tergal horns low, oblique, only slightly produced; inner dististyle with the outer basal lobe very large, produced into flattened blades; phallosome with the decurved apophyses expanded into broad flattened blades (fig. 159) . raysmithi Alexander 9 Wings conspicuously patterned with whitened and brown areas, the amount of dark color subequal to or exceeding the pale ground, the pale color including a narrow interrupted band in bases of all outer medial cells; male hypopygium with posterior border of eighth sternite produced into lateral lobes, their inner margin with a series of large flattened setae, the lobes smallest and with fewest such setae in vittatipennis, most elaborate in liastingsBe and polycantha where there are from seven to nine such structures (figs. 174-177) . 10 10 11 I1 vittatipennis group Wings pale, not or only slightly patterned with darker; male hypopygium with the eighth sternite not modified as described (figs. 121-123, 162, 163, 165-168, 170, 171, 178, 179) . . . . . . . 13 Darkened wing pattern more extensive, dark brown in hastingsae, paler in polycantha, involving most of cell R, including the basal fourth or more , . . 11 Darkened wing pattern basad of the cord more restricted, not including the broad bases of cells RandM . . . . . . , . . . . . . . 12 Wing pattern dark brown and white, the colors conspicuously contrasted; male hypopygium with the flattened modified setae of the eighth sternite fewer, about 6 or 7, narrower, tips acute; lobes of tergite obtuse; rods of ninth sternite strongly darkened, tips obtuse, phallosome with lateral apophyses large and powerful; inner dististyle long and narrow, posterior crest small (fig. 174) . hastingsae Alexander Wings with the brown pattern very extensive but paler and relatively inconspicuous, including most of cells R and M; male hypopygium with the flattened modified setae of eighth sternite about 8 or 9, broader, tips obtuse; lobes of tergite slightly pointed; rods of ninth sternite small and pale, pointed at tips; phallosome with lateral apophyses slender; inner dististyle with outer margin arched, polycantha Alexander posterior crest long and slender. (Extralimital: New Mexico, Arizona) 12 Large species (wing about 18 to 20 mm or more);
Page 3 and 4: THE CRANE FLIES OF CALIFORNIA
Page 5 and 6: I BULLETIN OF THE CALIFORNIA INSECT
Page 7 and 8: ! CONTENTS Introduction . . . . . .
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