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EXOTIC WOODY WEEDS Use of simulation models to predict future ...

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Due <strong>to</strong> its coarse geographic scale, the world distribution by country information can only be used for a<br />

coarse verification <strong>of</strong> the model. Ideally, the Australian distribution would only used for validation <strong>of</strong><br />

the model. However, because <strong>of</strong> the lack <strong>of</strong> knowledge <strong>of</strong> the exact climate preferences <strong>of</strong> rubber vine<br />

<strong>to</strong> some climate variables, it was necessary <strong>to</strong> include consideration <strong>of</strong> the Australian distribution <strong>of</strong> the<br />

plant in building the model. The use <strong>of</strong> reported physiological responses <strong>of</strong> the plant <strong>to</strong> weather<br />

conditions is complicated by the relationship between short-term weather variables and long-term<br />

climate averages.<br />

2.4.1 Present distribution <strong>of</strong> rubber vine<br />

2.4.1.1 Worldwide<br />

The global distribution <strong>of</strong> rubber vine is shown in Figure 2.6. It is primarily tropical, but extends in<strong>to</strong><br />

equa<strong>to</strong>rial and sub-tropical regions.<br />

Figure 2.6 Worldwide distribution <strong>of</strong> rubber vine (Jenkins 1944; Tomley 1995). Shaded areas indicate<br />

countries with rubber vine, not actual terri<strong>to</strong>ry occupied.<br />

2.4.1.2 Madagascar<br />

The climate <strong>of</strong> Madagascar in the natural range <strong>of</strong> rubber vine is characteristically dry tropical with a<br />

highly pronounced summer bias in the rainfall. It is important <strong>to</strong> note that Madagascar is a relatively<br />

small landmass, and has a limited amount <strong>of</strong> climatic variation. This means that rubber vine probably<br />

cannot fully express its climatic preferences. This limitation is most likely <strong>to</strong> be evident in the ability <strong>of</strong><br />

the plant <strong>to</strong> survive drier climates than are present in Madagascar because the end <strong>of</strong> the climatic<br />

gradient across the range <strong>of</strong> rubber vine is limited by sea.<br />

The existence <strong>of</strong> the congeneric Cryp<strong>to</strong>stegia madagascariensis Decne. within the northern range <strong>of</strong><br />

Cryp<strong>to</strong>stegia grandiflora provides a complication in the analysis. In Madagascar there is a distinct<br />

north-south rainfall gradient along the western coast. It is likely that C. madagascariensis is better able<br />

<strong>to</strong> <strong>to</strong>lerate the wetter conditions <strong>to</strong> the north, and outcompetes C. grandiflora. In the absence <strong>of</strong> this<br />

competition it is likely that C. grandiflora could persist in moister conditions than it has been noted<br />

occupying in Madagascar.<br />

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