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EXOTIC WOODY WEEDS Use of simulation models to predict future ...

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3.3 Prickly acacia population dynamics and management using the<br />

DYMEX model<br />

A vast number <strong>of</strong> model <strong>simulation</strong>s have been run and only representative results are presented here <strong>to</strong><br />

illustrate the nature <strong>of</strong> the <strong>simulation</strong>s and results. Most model runs were carried out using weather<br />

data sequences and paddock conditions representative <strong>of</strong> Marathon Station, near Hughenden (143º 34'E<br />

20º 54' S). The modelled patterns <strong>of</strong> phenology, germination, individual development, survival,<br />

competition, and self-thinning are both qualitatively and quantitatively consistent with field studies.<br />

Model runs have been limited <strong>to</strong> 40 year periods (approximately the lifespan <strong>of</strong> prickly acacia) due <strong>to</strong><br />

available weather data and computation power. Over such short runs, the model is sensitive <strong>to</strong> starting<br />

conditions. This is consistent with field observations <strong>of</strong> similar aged invasions that show marked<br />

differences in infestation density and population structure that can not be attributed <strong>to</strong> biophysical or<br />

land use differences, suggesting invasion his<strong>to</strong>ry as the proximal cause.<br />

3.3.1 Impacts <strong>of</strong> prickly acacia invasion<br />

In order <strong>to</strong> estimate the effects <strong>of</strong> invasion by prickly acacia on pasture productivity, the relationship<br />

between the density <strong>of</strong> prickly acacia and pasture production (Mooy et al. 1992) was used in a<br />

<strong>simulation</strong> <strong>of</strong> the effect <strong>of</strong> planting trees along a bore drain on a cattle property. The results (Figure<br />

3.20) show the dramatic decline in pasture production after 40 years (only the last 30 years <strong>of</strong> the<br />

<strong>simulation</strong> are shown).<br />

Basal Area Density (m 2 /ha)<br />

5<br />

4.5<br />

4<br />

3.5<br />

3<br />

2.5<br />

2<br />

1.5<br />

1<br />

0.5<br />

0<br />

2000<br />

1800<br />

1600<br />

1400<br />

1200<br />

1000<br />

800<br />

600<br />

400<br />

200<br />

0<br />

Pasture Production Density (kg/ha)<br />

Upland Basal<br />

Area Density<br />

(m^2 ha^-1)<br />

1957<br />

1959<br />

1961<br />

1963<br />

1965<br />

1967<br />

1969<br />

1972<br />

1974<br />

1976<br />

1978<br />

1980<br />

1982<br />

1985<br />

1987<br />

1989<br />

1991<br />

Upland Pasture<br />

Production<br />

Density (kg ha^-<br />

1)<br />

Figure 3.20 Simulated impact <strong>of</strong> increasing prickly acacia basal area density on pasture production at<br />

Hughenden, north Queensland. Pasture production estimates were derived using a regression equation<br />

from Mooy et al. (1992).<br />

3.3.2 Effects <strong>of</strong> climate change on prickly acacia population dynamics<br />

In Australia, the projected effect <strong>of</strong> increases in temperature and water use efficiency is <strong>to</strong> intensify the<br />

population dynamics processes <strong>of</strong> prickly acacia. The consequences <strong>of</strong> this are <strong>to</strong> increase the rate <strong>of</strong><br />

invasion <strong>of</strong> uplands, and <strong>to</strong> increase the amount <strong>of</strong> prickly acacia biomass that can be sustained,<br />

particularly in uplands. It is important <strong>to</strong> note that due <strong>to</strong> self-thinning processes (intra-specific<br />

competition) and the law <strong>of</strong> constant yield, stem density can vary widely for a given biomass density.<br />

Thus stem density is not a good indica<strong>to</strong>r <strong>of</strong> the state <strong>of</strong> a paddock with respect <strong>to</strong> weediness. Biomass<br />

density <strong>of</strong> woody plants is <strong>of</strong> interest <strong>to</strong> climate change research because it is related <strong>to</strong> the amount <strong>of</strong><br />

carbon sequestered in the tree and shrub material. It is also closely related <strong>to</strong> basal area density, which<br />

in turn is related <strong>to</strong> pasture production. Unfortunately, neither <strong>of</strong> these measures is easily perceived<br />

through casual inspection. Canopy cover <strong>of</strong> prickly acacia ie proportion <strong>of</strong> the paddock covered by the<br />

31

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