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~ G0UNmB¥ BER0BmS 15<br />
Current genetic conservation activities in situ and ex situ<br />
Since the early 1980s, gene reserves have been designated as follows: 6789.1 ha for Q. robur,<br />
265.2 ha for Q. petraea and 2820.4 ha for F. sylvatica. The distribution of gene reserves in<br />
Ukraine is given in Table 1. The largest reserve areas are concentrated in forest regions.<br />
Presently the inventory of gene reserves is being undertaken. This will provide the<br />
opportunity to estimate the current situation of genetic resources.<br />
For several years, studies on European beech diversity in the Carpathian region and<br />
adjacent territories have been carried out by the Ukrainian State University of Forestry and<br />
Wood Technology (Lviv), together with Technical University in Zvolen, Slovakia. Genetic<br />
diversity and differentiation of beech populations (20 European beech and 7 Crimean beech)<br />
have been studied by electrophoresis.<br />
Genetic differentiation was estimated on the basis of genetic distances. The results<br />
confirm a low degree of differentiation within F. sylvatica compared with F. orientalis, the<br />
values of subpopulation differentiation of European beech populations being only<br />
approximately 30% of the differentiation among F. oriental is populations. Within European<br />
beech populations from western Ukraine a slight differentiation between populations from<br />
the southwestern and northeastern macroslopes of the Carpathians, and plain populations<br />
from the northeastern limit of beech distribution range was observed. Crimean beech is also<br />
much more differentiated than Carpathian beech. Crimean beech occupies an eccentric<br />
position between both beech species; however, it is much more similar to F. orientalis than to<br />
F. sylvatica (Paule et al. 1993; Vysny et al. 1995; Gomory et al. 1998a, 1998b).<br />
The intensity of reproduction in beech root meristem cells was examined in cytological<br />
studies by the University together with the Institute of Forestry and Forest Melioration. The<br />
results led to the conclusion that some differences exist in the mitotic activity of root<br />
meristem cells of beech seedlings from different populations. Since the mitotic kinetics of<br />
the meristematic cells of all plants depend not only on ext~rnal factors but also on internal<br />
factors, it is quite possible that the results reflect the genetic diversity between populations<br />
(Kyrychenko et al. 1995).<br />
Clonal propagation of European beech and sessile oak is studied at the Laboratory of<br />
Tissue Culture. For this purpose explants were taken from different vegetative and<br />
generative plant parts of beech (buds of different-aged trees, hypocotyls, cotyledons)<br />
(Bazyuk and Fedyaj 1995).<br />
Relevant nature protection policies and activities<br />
Among the legislative acts related to conservation of genetic resources, the Law on Natural<br />
Reserves of Ukraine (of 5 May 1993) is in force. According to this law any activity which<br />
might affect negatively the state of protected areas is forbidden. The Committee of<br />
Environmental Protection is the administrative unit specialized in this field. The protection<br />
is implemented by the institutions on the territory in which they are situated. Protection of<br />
forest areas is carried out by regional forest offices.<br />
Table 1. Gene reserves of Social Broadleaves in Ukraine<br />
Quercus robur Quercus eetraea Faflus syJvatica<br />
Natural zone Area {ha} Number Area {ha} Number Area (ha) Number<br />
Mixed forest 2526.3 80 52.4 1 2.0 1<br />
Forest-steppe 3935.8 160 29.0 2 1789.8 60<br />
Steppe 269.0 1 128.0 7 0.0 0<br />
Carpathians 58.0 1 22.1 2 1028.6 7<br />
Crimea 0.0 0 33.7 4 0.0 0<br />
Total 6789.1 257 265.2 16 2820.4 68