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Kernel Home Range Estimation for ArcGIS, using VBA - Fish and ...

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a b c<br />

Figure 4.5.3.3. Inclusion of points in a density estimate based on the truncated search area when<br />

<strong>using</strong> the covariance bias method of st<strong>and</strong>ardization.<br />

It should be noted that this method is not recommended. It is provided as an option in ABODE<br />

should the user wish to make comparisons between home range estimators that may only<br />

provide this method of st<strong>and</strong>ardization. This method is heavily biased by not incorporating the<br />

covariance between x <strong>and</strong> y.<br />

5. <strong>Home</strong> <strong>Range</strong> Asymptotes<br />

5.1. Why we should look at them<br />

Harris et al. (1990) suggested that a home range analysis should be done <strong>using</strong> data that<br />

encompass the full range of variation in movement behavior attributable to sex <strong>and</strong> age<br />

differences. This is only possible if a representative sample (generally evenly spaced in time) is<br />

obtained <strong>for</strong> the entire sampling duration. To ensure that the sampling duration covers the full<br />

range of behavior exhibited by the animal, home range asymptotes are necessary. This should<br />

typically be done <strong>using</strong> a preliminary dataset, be<strong>for</strong>e the majority of the data are collected. The<br />

point in time (conversely the number of locations required) where the home range reaches an<br />

asymptote will indicate what the sampling duration (sample size) requirement should be. If home<br />

ranges do not asymptote, then the user may not have a representative sampling duration <strong>for</strong> the<br />

time period. Alternatively, a lack of asymptote may indicate a multiscaled home range<br />

(Gautestad <strong>and</strong> Mysterud, 1993). Harris et al. (1990) suggested the use of “area observation<br />

plots” (Otis <strong>and</strong> White, 1999) to determine the number of locations required to obtain a stable<br />

estimate of home range size (Stickel, 1954; Hawes, 1977). Gautestad <strong>and</strong> Mysterud (1995)<br />

proposed alternatively that home ranges are not asymptotic, but rather increase according to the<br />

power law (square root of the number of locations). This may be true <strong>for</strong> MCP analyses, but<br />

kernel estimators are relatively robust towards sample size issues (Seaman et al. 1999). Using<br />

simulation data, Seaman et al. (1999) showed that kernels gave stable estimates at about 50<br />

30

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