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full text - Caspar Bgsu - Bowling Green State University

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2. Evolutionary Aspects of Aggression: The Importance of Sexual Selection 15<br />

the benefit of winning—the value of the contested item—is much larger than the<br />

cost of losing. If the choice is reproduction or death, fights become deadly. This is<br />

why fights among elephant seals are so fierce and bloody. The chance to mate<br />

occurs only once per year and most males never even get close. For the successful<br />

males it is another story—in a study of Southern elephant seals, harem holders<br />

accounted for 89.6% of the recorded paternities (Fabiani et al., 2004; Fig. 2.2).<br />

The sequential assessment game is a variant of a game theoretical setup<br />

termed the “hawk-dove game” (see also Chapter 3). In this game, there are two<br />

possible strategies: always fight (“hawk”) and always yield (“dove”), where it is<br />

assumed that the two competitors have equal fighting ability. An Evolutionarily<br />

Number of males Number of males<br />

12<br />

8<br />

4<br />

0<br />

12<br />

8<br />

4<br />

RUB96<br />

SF96<br />

SF97<br />

0 1 2 7 0 1 12 1 24<br />

SI196 SI296 SI297<br />

0<br />

0 1 24 0 1 2 8 21 0<br />

12<br />

1 2 8 32<br />

Number of males<br />

8<br />

4<br />

SM96<br />

HH<br />

NHH<br />

0<br />

0 2 12<br />

Number of paternities<br />

Figure 2.2. Number of paternities achieved by the harem holder (HH) and the other males (NHM)<br />

associated with each harem in seven different populations of Southern elephant seals<br />

(from Fabiani et al., 2004).

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