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LILIACEAE - China

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马 甲 菝 葜 ma jia ba qiaVines climbing. Stem branched, terete, 1–2 m, woody;branchlets occasionally zigzagged. Petiole 1–2(–2.5) cm, narrowlywinged for 1/5–1/4 its length; abscission zone at middle;tendrils usually present. Leaf blade lanceolate to ovate-oblong,6–17 × 2–8 cm. Inflorescence of 1(or 2) umbels, basally prophyllate;peduncle 1–1.5 cm, proximally articulate; umbels ofboth sexes densely 20–30-flowered, base slightly thickened.Male flowers: tepals yellowish green, 3–4.5 × ca. 1 mm; stamens3–4 mm. Female flowers: tepals 1.5–2 × ca. 0.6 mm; staminodes6. Berries yellowish red to black, globose, 6–7 mm indiam. Fl. Sep–Mar, fr. Oct–Nov.Forests, forest margins, thickets, shaded places on slopes; 100–2800 m. Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hubei, Hunan,Jiangxi, Sichuan, Taiwan, Yunnan, Zhejiang [Bhutan, Cambodia,NE India, Indonesia, Laos, Malaysia, Myanmar, Philippines, Sikkim,Thailand, Vietnam].This species is very difficult to treat taxonomically due to theextreme polymorphism of the included entities.1a. Main veins obviously concave on adaxialleaf surface ................................... 52e. var. impressinervia1b. Main veins raised on adaxial leaf surface(sometimes midvein slightly concave).2a. Leaf blade usually ovate or ovate-lanceolate,usually 1–3 × as long as wide.3a. Leaf blade usually papery, adaxiallynot or slightly shiny; peduncle usuallyshorter than petiole ............. 52a. var. lanceifolia3b. Leaf blade usually leathery, adaxiallyshiny; peduncle usually longer thanpetiole ........................................ 52b. var. opaca2b. Leaf blade lanceolate to narrowly oblonglanceolate,usually 5–7 × as long as wide.4a. Branchlets ± zigzagged; pedunclelonger than petiole; tepals yellowishgreen ...................................... 52c. var. elongata4b. Branchlets not zigzagged; peduncleshorter than or subequaling petiole;tepals purple ....................... 52d. var. lanceolata52a. Smilax lanceifolia var. lanceifolia马 甲 菝 葜 ( 原 变 种 ) ma jia ba qia (yuan bian zhong)Smilax micropodaA. de Candolle.Leaf blade usually ovate to ovate-lanceolate, usually 1–3 ×as long as wide, papery, adaxially not or slightly shiny, mainveins raised adaxially (sometimes midvein slightly concave). 2n= 32.Forests, thickets, shaded places on slopes; 600–2800 m. Guangxi,Guizhou, Hubei, Sichuan, Yunnan [Bhutan, NE India, Laos, Myanmar,Sikkim, Thailand, Vietnam].52b. Smilax lanceifolia var. opaca A. de Candolle in A. deCandolle & C. de Candolle, Monogr. Phan. 1: 57. 1878.暗 色 菝 葜 an se ba qiaSmilax laevis Wallich ex A. de Candolle; S. laevis var.ophirensis A. de Candolle; S. laevis var. parkii A. de Candolle;S. lanceifolia subsp. opaca (A. de Candolle) T. Koyama; S.opaca (A. de Candolle) J. B. Norton.Leaf blade leathery, usually ovate to ovate-lanceolate, usually1–3 × as long as wide, adaxially obviously shiny, mainveins raised adaxially (sometimes midvein slightly concave).Forests, thickets, shaded places on slopes; 100–1000(–2000) m.Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hunan, Jiangxi, Taiwan,Yunnan, Zhejiang [Cambodia, Indonesia, Laos, Malaysia, Thailand,Vietnam].52c. Smilax lanceifolia var. elongata (Warburg) F. T. Wang &Tang, Fl. Reipubl. Popularis Sin. 15: 220. 1978.折 枝 菝 葜 zhe zhi ba qiaSmilax microphylla C. H. Wright var. elongata Warburg inDiels, Bot. Jahrb. Syst. 29: 259. 1900; S. austrosinensis F. T.Wang & Tang; S. tortipetiolata H. Léveillé & Vaniot.Branchlets zigzagged. Leaf blade lanceolate or narrowlyoblong-lanceolate, usually 5–6 × as long as wide, thickly paperyto leathery, main veins raised adaxially (sometimes midveinslightly concave). Peduncle longer than petiole. Tepals yellowishgreen.Forests, shaded places on slopes; 500–2000 m. Guangdong,Guangxi, Guizhou, Jiangxi, Sichuan, ?Yunnan, Zhejiang.52d. Smilax lanceifolia var. lanceolata (J. B. Norton) T. Koyama,Quart. J. Taiwan Mus. 13: 26. 1960.长 叶 菝 葜 chang ye ba qiaSmilax cocculoides Warburg var. lanceolata J. B. Nortonin Sargent, Pl. Wilson. 3: 11. 1916.Leaf blade lanceolate to narrowly oblong-lanceolate, usually5–7 × as long as wide, main veins raised adaxially (sometimesmidvein slightly concave). Peduncle shorter than or subequalingpetiole. Tepals purple.Forests, forest margins; 1500–2000 m. S Yunnan.52e. Smilax lanceifolia var. impressinervia (F. T. Wang &Tang) T. Koyama, Quart. J. Taiwan Mus. 13: 26. 1960.凹 脉 菝 葜 ao mai ba qiaSmilax impressinervia F. T. Wang & Tang, Sinensia 5: 425.1934.Leaf blade lanceolate to oblong-lanceolate, thinly leathery;main veins 3, obviously concave adaxially. Peduncle subequalingpetiole.Shaded places in forests; 1000–2000 m. Guangxi, Guizhou,Yunnan.53. Smilax astrosperma F. T. Wang & Tang, Fl. Hainan. 4: 534.1977.灰 叶 菝 葜 hui ye ba qiaVines climbing. Stem branched, terete, 1–2 m, woody,sometimes sparsely prickly. Petiole 1–1.5 cm, narrowly wingedfor 1/5–1/4 its length; abscission zone at or above middle;

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