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248 R.M. Ros, V. Mazimpaka et al.[404] Muñoz (1998) did not accept Grimmia sessitana De Not. and considered it togetherwith its synonyms (G. alpestris var. sessitana (De Not.) I. Hagen, G. anceps Boulay,G. sessitana var. subsulcata Limpr., G. subcaespiticia Schiffn., Grimmia subsulcataLimpr., and Grimmia tenerrima Renauld & Cardot) within the concept ofG. reflexidens Müll. Hal. This criterion was adopted in Hill et al. (2006) and also inthis checklist, altough Greven (2011) considers G. sessitana a distinct species that ispresent throughout the study area in Andorra, Bulgaria, Corsica, Italy, Sardinia,and Spain.[405] The proposal of Ochyra et al. (2003) of segregating the Campylium hispidulumcomplex (represented in the Mediterranean region by C. calcareum andC. sommerfeltii) from the genus Campylophyllum (Hypnaceae) (represented in theMediterranean by C. halleri) and including the species of the mentioned complex inthe genus Campylidium (Amblystegiaceae) has received molecular support in thestudies of Gardiner et al. (2005) and Ignatov et al. (2007). Therefore this treatment ishere followed.[406] Didymodon verbanus (W.E. Nicholson & Dixon) Loeske (= Eucladium verbanumW.E. Nicholson & Dixon), known in the studied area from northern Italy, was treatedby Jiménez (2006b) as a synonym of D. glaucus. This criterion was followed in Hillet al. (2006) and also here, although Kuªera (2000) considered it a different species.[407] All occurrences of the epithet “durieui”, and combinations based on, need to bechanged to “duriaei”, because this is the original spelling and it does not seem to be anorthographic error that would have been corrected in the sense of Art. 60.1 and60.3 of ICBN.[408] The molecular studies of Gardiner et al. (2005) and Ignatov et al. (2007) onLeskeaceae s.l., suggest that the genera Pseudoleskea and Ptychodium, that aresometimes separated from Lescuraea should be placed within the latter genus andexcluded from Leskeaceae. Of the species present in the Mediterranean, onlyPseudoleskea artariae (= Pseudoleskeopsis artariae (Thér.) Thér.) has not beenstudied at molecular level, therefore its generic assignement is still uncertain and ithas been maintained in this paper in Pseudoleskea as Hill et al. (2006) proposed.[409] According to Vanderpoorten et al. (2003) Leucobrym albidum and L. juniperoideumshould be treated as synonyms, although this was not followed by Hill et al. (2006).[410] Bell & Hyvönen (2010) observed in a phylogenetic study that the species ofPolytrichastrum sect. Aporotheca (represented in the Mediterranean area byP. formosum, P. longisetum and P. pallidisetum) should be reinstated in Polytrichum.[411] According to a molecular and morphological study carried out by Hedenäs &Rosborg (2008), the Pseudocalliergon species should be moved to Drepanocladus.[412] Hedenäs (2006) transferred the dioicous species of Warnstorfia (represented in theMediterranean by W. exannulata and W. sarmentosa) to Sarmentypnum based onthe results of that paper and those previously published by Hedenäs et al. (2005).[413] Anoectangium angustifolium Mitt. (= Anoectangium compactum var. madeirenseGeh., Anoectangium madeirense Schimp. nom. inval. in synon.) reported from theAzores, Canary Islands, France, and Madeira was synonymized with A. aestivum byDirkse et al. (1993) based on the overlapping observed between both species.Nevertheless, recent unpublished molecular data support the distinction of bothspecies (Kuªera, pers. comm.).[414] Barbula bescherellei Sauerb. was reported from Egypt in the unpublished Master ofScience Thesis of Mansi (1988) and compiled by El-Saadawi et al. (1999). As thistaxon was described from America, its occurrence in Egypt is unlikely. It wassynonymized with Didymodon rigidulus by Zander (1981), but the concept ofthis author at that time was extremely broad and does not correspond with thepresent one.
Mosses of the Mediterranean, an annotated checklist 249[415] Grimmia limprichtii Kern was considered by Muñoz & Pando (2000) a synonym ofGrimmia anodon Bruch & Schimp. and this has been the criterion followed here,although Maier & Geissler (1995) and Greven (2009b) synonymized it withG. tergestina.[416] According to Greven (2011) Grimmia crinitoleucophaea Cardot should be the correctname for G. poecilostoma Cardot & Sebille ex Limpr. The proposal of Muñoz (2000)to conserve the last one was rejected by Zijlstra (2002), as this name seems to besuperfluous and invalid.[417] The European endemic Grimmia pilosissima Herzog was considered a synonym ofGrimmia montana by Muñoz & Pando (2000) and this has been the criterion followedhere. However, Greven (1994b) considered it a distinct species.Maier (2010) considered G. ungeri to be a synonym of G. montana, but the two taxaare treated here separately.[418] Tortula densa (Velen.) J.-P. Frahm was considered by Hill et al. (2006) to be asynonym of Syntrichia calcicola, althoug Kramer (1980) had a different opinion andsynonymized it with S. ruralis s.s.[419] Amblystegium serpens was reported from Crete (as A. juratzkanum) by Baumgartner(1943). The later record of A. juratzkanum in Düll (1966) is regarded as unconfirmed(“unbestätigt”) by Düll (1979).[420] Schistidium helveticum is reported from Crete as S. singarense without an indicationof date by Blom (1996), based on a specimen collected by K. Rechinger, probably in1942. Schistidium brunnescens subsp. brunnescens is reported from Greece by Blom(1996), based on a specimen collected by C. Haussknecht in 1885.[421] Pleuridium acuminatum is reported from Cyprus by Townsend (1965), based on aspecimen collected in 1957. Ptychostomum pseudotriquetrum var. bimum wasreported from Cyprus also by Townsend (1965), based on a specimen collected byE.W. Kennedy in 1937 and reported by Taylor (1952) as B. obconicum.[422] The occurrence in Spain of Cleistocarpidium palustre (Simó, 1977), is based an oldcollection of Durieui de Maisonneuve.[423] Brachythecium velutinum var. athoum Podp. was described from Greece but itsidentity has not been revised since. If it merits recognition, then a new combination inthe genus Brachytheciastrum Ignatov & Huttunen may be required.[424] Bryum capillare var. cavifolium Breidl. was described from Greece but its identity hasnot been revised since. If it merits recognition, then a new combination in the genusPtychostomum Hornsch. may be required.[425] Funaria mediterranea var. sinistra Schiffn. was described from Greece but its identityhas not been revised since. If it merits recognition, then a new combination in thegenus Entosthodon Schwägr. may be required.Acknowledgements. We thank several bryologists who have helped us to solvesome taxonomic uncertainties or have provided literature references. Among them specialthanks are addressed to Lars Hedenäs, David T. Holyoak, Denis Lamy, Marshal R. Crosbyand Jesús Muñoz. We also thank Mahmoud Magdy (University of Murcia) for his help withcomputer resources and issues and Jan Kuªera for the detailed and very valuable correctionof a previous version of the manuscript. One of the co-authors, Tom Blockeel, made anadditional contribution by revising the English language of the manuscript. This work hasbeen carried out with financial support from the Spanish Ministry of Science andInnovation (Projects CGL2008-00275/BOS and CGL2011-22936/BOS) and co-funded withFEDER program from the EU.
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