Mosses of the Mediterranean, an annotated checklist - Optima-bot.org

226 R.M. Ros, V. Mazimpaka et al.[94] Tortula wilczekii Meyl. was described from Algeria as Tortula mairei Meyl. nom.illeg., a later homonym. However, its identity has not been revised because the typehas not been traced (Ros, unpublished).[95] Tortula buyssonii (H. Philib.) Broth. was described from France as Barbula buysonniiH. Philib., but its identity has not been revised.[96] Barbula montenegrina Breidl. & Szyszyl., often considered a synonym of Tortulalingulata Lindb., has been synonymised by Ko≠nar & Kuªera (2010) with T. muralisvar. obtusifolia. As this is the only report of Tortula lingulata in the Mediterraneancountries, it should be excluded from the area.[97] The species included in the genus Phascum by Hill et al. (2006), Phascum cuspidatumHedw. and Phascum vlassovii Laz., are here included in the genus Tortula as Tortulaacaulon and Tortula vlassovii following the proposals of Zander (1993) and Werneret al. (2002, 2004) for the former and Ros & Herrnstadt (2010) for the latter. Also Ros& Herrnstadt (2010) reduced Phascum galilaeum Herrnst. & Heyn. to a variety ofTortula acaulon and synonymised Pottia gemmifera Herrnst. & Heyn., also fromIsrael, with Tortula caucasica.[98] The genus Protobryum is accepted in Hill et al. (2006) for the single speciesP. bryoides (Dicks.) J. Guerra & M.J. Cano. We include this species in the genusTortula as proposed by Zander (1993) and corroborated subsequently from moleculardata by Werner et al. (2002, 2004).[99] According to Ros et al. (2008), the names Tortula caucasica Broth. and T. lindbergiishould replace T. modica R.H. Zander and T. lanceola R.H. Zander respectively.[100] Tortula kneuckeri Broth. & Geh. was considered a nomen nudum by Wijk et al. (1969)based on Brotherus (1924), who did not include a species description or illustration.However, a description of this species had been given earlier by Geheeb (1903), whichsubstantiates its recognition. More recently it has been collected and described fromSinai (Egypt) by Abou-Salama (2001).[101] Tortula mucronifera was described from the Arabian Peninsula and Jordan by Freyet al. (1994) and later reported from Morocco by Cano et al. (2002).[102] Ko≠nar & Kolá÷ (2009) in a biosystematic study of the Tortula muralis complex usingmultivariate analysis of morphological characters, a cultivation experiment andcytological screening showed that subsp. obtusifolia (= T. obtusifolia (Schwägr.)Mathieu) was invariably haploid, but that both haploid and diploid plants were shownto occur in subsp. muralis. The pattern of morphological variability however allowedthe recognition of the two subspecies within T. muralis and two varieties within subsp.muralis. In the case of var. aestiva, they maintained the varietal rank. Even though ithad been synonymised with the type variety by Cano (2006a), Ko≠nar & Kolá÷ (2009)observed that the main character of the gametophyte used for the determination ofthis variety (costa excurrency) proved to be stable in cultivation, indicating that thischaracter is under genetic control.[103] Tortula plinthobia (Sull. & Lesq.) Broth. was reported from Egypt by Refai (2001)but this is the only Mediterranean report of this species known from North America.This single report requires verification.[104] Triquetrella papillata (Hook. f. & Wilson) Broth. was reported from Egypt by Badawiet al. (1987) and El-Saadawi et al. (1987), but later considered a misidentification byEl-Saadawi et al. (1999). It is therefore rejected from the area.[105] Weissia leptocarpa Schimp. ex Besch. hom. illeg. is a doubtful and illegitimate namethat was described from Portugal (Levier 1880), and later reported from Turkey(Wettstein, 1889) and Algeria (Jelenc, 1955), and later included in some of thecompilations of these countries. The identity of the name has not been revisedrecently.