Gyraulus parvus (Mollusca: Gastropoda) in the Czech Republic

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Acta Soc. Zool. Bohem. 66: 141–150, 2002ISSN 1211-376XNew Tertiary dragonflies from Lower Oligocene of the eské stedohoíMts and Lower Miocene of the Most Basin in the Czech Republic(Odonata: Anisoptera)Jakub PROKOP 1, 2) & André NEL 3)1)Department of Zoology, Charles University, Vininá 7, CZ–128 44, Praha 2, Czech Republic;e-mail: jprokop@natur.cuni.cz2)Department of Palaeontology, Charles University, Albertov 6, CZ–128 43 Praha 2, Czech Republic3)Laboratoire d’Entomologie et CNRS UMR 8569, Museum national d’Histoire naturelle, 45 rue Buffon,F–75005 Paris, France; e-mail: anel@mnhn.frReceived January 30, 2002; accepted March 23, 2002Published June 28, 2002Abstract. Two new representatives of the clade Aeshnoptera are described from the Lower Oligocene andLower Miocene of northern Bohemia (Czech Republic), i. e. Kvacekia infuscata gen. n. et sp. n. (Aeshnidae)and Gomphaeschna miocenica sp. n. (Gomphaeschnidae). Kvacekia gen. n. seems to be closely related tothe Cenozoic genus Oligaeschna Piton et Théobald, 1939 and the recent genus Oplonaeschna Selys, 1883.Gomphaeschna miocenica sp. n. wing venation has particular wing coloration and distinctly differentcharacters from all previously described species of the genus. A holarctic distribution in fossil history isproposed for both Oplonaeschninae and Gomphaeschninae.Taxonomy, fossil, description, Insecta, Anisoptera, Odonata, Aeshnoptera, Gomphaeschnidae,Aeshnidae, Kvacekia infuscata gen. n. et sp. n., Gomphaeschna miocenica sp. n., Tertiary, LowerOligocene, Lower Miocene, Central Europe, Palaearctic regionINTRODUCTIONTertiary fossil dragonflies have been recorded from several sites such as Pochlovice near Kynšperk,Sokolov, Jehliná near Sokolov belonging to Cypris Shale and from Bílina mine of Most Formationin northwestern Bohemia (Bachmayer 1952, Heer 1849, Handlirsch 1907, Kukalová 1962, íha 1977,1979, Prokop & Nel 2000). Up till now all these recorded fossils, both larvae and adults, belong toLibellulidae and Aeshnidae (Prokop 2002).Cenozoic insect fauna of northwestern Bohemia is preserved in fluvio-lacustrine deposits ofthe Krušné hory piedmont basins and the eské stedohoí Mts. About 16 localities representingseveral different palaleoenvironments dated from Upper Eocene to Lower Miocene are investigated.The relatively diverse insect fauna from Lower Oligocene (Ruppelian/Chattian) of Kundraticenear Litomice is preserved in several interbeds of the brownish diatomite of Ústí Formation (Fig.1b). The site is well known because of studied palaeobotanical and palaeoichthyological records(Kvaek & Walther 1998, Obrhelová 1979). Another famous site is the Bílina mine (the formerMaxim Gorkij mine) situated in the Most Basin near the town of Bílina (Fig. 1b). The stratigraficalattribution is to the Most Formation of the Lower Miocene (Eggenburgian/Ottnangian) and insectsare preserved in three fossiliferous horizons (Clayey Superseam Horizon, Delta Sandy Horizon,Lake Clayey Horizon) (Prokop 2002). The site has been observed from other paleoscientificaspects as palaeobotany, palynology and sedimentology (Bžek et al. 1992, Dašková 2000, Kvaek1998, Rajchl & Uliný 1999, Sakala 2000).141
In the following study we use the wing venation nomenclature of Riek & Kukalová-Peck (1984),amended by Kukalová-Peck (1991), Nel et al. (1993), Bechly (1996) and Bechly et al. (2001). Wefollow the phylogenetic classification of Anisoptera proposed by Bechly et al. (1996, 2002).SYSTEMATIC PALAEONTOLOGYAeshnoptera Bechly, 1996Family Aeshnidae Leach, 1815Genus Kvacekia gen. n.TYPE SPECIES. Kvacekia infuscata sp. n.DIAGNOSIS. The new genus very similar and probably closely related to recent genus OplonaeschnaSelys, 1883 and fossil genus Oligaeschna Piton et Théobald, 1939. It differs from these generaby the following combination of characters (fore wing structures): (1) pterostigma elongate, covering5–6 cells; (2) cubito-anal area very broad, with more than 9 rows of cells between CuA andposterior wing margin (possible autapomorphy, but character unknown in some Oligaeschnaspecies); (3) 5 rows of cells in area between IR2 and RP2; (4) hypertriangle with 4 or more crossveins;(5) wing dark infuscate; (6) about 18–20 postnodal cross-veins (14 to 16 in the Oplonaeschnaand Oligaeschna species, but character state unknown in Oligaeschna lapidaria). Characters(2) and (5) are probably autapomorphies of Kvacekia gen. n., but no relative autapomorphy ofOligaeschna is known. Only the discovery of better-preserved specimens of both taxa will permitto solve the problem of the possible paraphyly of Oligaeschna relatively to Kvacekia gen. n.ETYMOLOGY. Kvacekia gen. n. (feminine in gender), named in honour of Prof. Zlatko Kvaek,palaeobotanist from Charles University in Prague.Kvacekia infuscata sp. n.(Figs 2, 3)DIAGNOSIS. That of the genus.DESCRIPTION. A nearly complete fore wing, broken in nodal region and with part basal of arculusmissing; whole wing surface dark fuscous with some lighter zones (see Fig. 2); fore wing, about52 mm long (assumed from fragment) and 12 mm wide; distance from estimate base to nodus, about25 mm; distance from nodus to wing apex, about 27 mm; nodus nearly midway between base andapex; distance from nodus to pterostigma, 12.4 mm; distance from pterostigma to apex, 5.6 mm;pterostigma rather long, 5.3 mm long and 0.9 mm wide, covering approximately 5–6 cells; pterostigmalbrace slightly obliquely aligned with proximal side of pterostigma; 19 visible antenodal crossveinsof first row between C and ScP not aligned with the 16 visible corresponding antenodalcross-veins of second row between ScP and RA; 18 visible postnodal cross-veins, not well alignedwith 19 visible subpostnodal cross-veins, maybe 1–2 missing postnodal cross-veins close tonodus; more than 12 secondary antenodal cross-veins visible between Ax2 and nodus; more than4 antenodal cross-veins of second row between Ax2 and Ax1; Ax1 being 6.2 mm and Ax2 12.9 mmfrom wing base; hypertriangle crossed by 4–5 cross-veins; arculus, anal area, median and submedianspace, and subdiscoidal triangle not preserved; discoidal triangle elongate and divided into7 smaller cells, its costal side being 9.2 mm long, distal side 8.3 mm long and proximal side 3.1 mmlong; width of postdiscoidal area just behind discoidal triangle, 4 mm, probable width along posteriorwing margin, 11.4 mm; 3 rows of cells in postdiscoidal area just distal of discoidal triangle; ashort convex supplementary sector (trigonal planate) in postdiscoidal area, beginning one cell142
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Acta Soc. Zool. Bohem. 66: 81-84, 2
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REFERENCESBERAN L. & HORSÁK M. 199
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Tab. 1. Nematodes studied with the
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Tab. 2. The dimensions (mm) of thir
Page 14 and 15: Family Acuariidae Railliet, Henry e
Page 16 and 17: Fig. 7. Cosmocephalus obvelatus (Cr
Page 18 and 19: A c k n o w l e d g e m e n t sThe
Page 20 and 21: Acta Soc. Zool. Bohem. 66: 99-119,
Page 22 and 23: Figs 1-10. Harpalus (Harpalus) atra
Page 24 and 25: Figs 11-19. Harpalus (Harpalus) cis
Page 26 and 27: Figs 20-27. Harpalus (Harpalus) lut
Page 28 and 29: Figs 28-35. Harpalus (Harpalus) pic
Page 30 and 31: Figs 36-44. Harpalus (Harpalus) sax
Page 32 and 33: Figs 45-53. Harpalus (Harpalus) ser
Page 34 and 35: Figs 54-61. Harpalus (Harpalus) sol
Page 36 and 37: Figs 62-70. Harpalus (Harpalus) xan
Page 38 and 39: Based on the study of our larval ma
Page 40 and 41: EMDEN F. I. van 1942: A key to the
Page 44 and 45: and D. papillifera (Fuhrmann, 1908)
Page 46 and 47: about 1 mm) with only a few eggs in
Page 48 and 49: The number of catfish examined each
Page 50 and 51: number (3.65) of trematodes with ma
Page 52 and 53: cestode was distinctly associated w
Page 54 and 55: from individual months. Tab. 2 show
Page 56 and 57: number (5.57) of males occurred in
Page 58 and 59: intermediate host, copepod, catfish
Page 60 and 61: CHUBB J. C. 1982: Seasonal occurren
Page 64 and 65: efore distal angle of discoidal tri
Page 66 and 67: cells between them and IR2 or MA”
Page 68 and 69: character is less pronounced in Oli
Page 70 and 71: REFERENCESBACHMAYER F. 1952: Fossil
Page 74 and 75: elow) and a 170×90×70 type was us
Page 76 and 77: the brood, first into heaps near th
Page 78 and 79: trophallaxis until the termination
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Gyraulus parvus (Mollusca: Gastropoda) in the Czech Republic

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