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Maize: Origin, Domestication, and its Role in the Development of Culture

by Duccio Bonavia

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24<br />

<strong>Maize</strong>: <strong>Orig<strong>in</strong></strong>, <strong>Domestication</strong>, <strong>and</strong> Its <strong>Role</strong> <strong>in</strong> <strong>the</strong> <strong>Development</strong> <strong>of</strong> <strong>Culture</strong><br />

Teos<strong>in</strong>te<br />

In his 1893 study, Harshberger accurately described <strong>the</strong> area where annual teos<strong>in</strong>te<br />

grows, as well as <strong>the</strong> archaeological zones where it grew <strong>in</strong> <strong>the</strong> wild state<br />

(Wilkes, 1989: 446). It is, however, worth recall<strong>in</strong>g that this plant was not<br />

studied <strong>in</strong> depth until Wilkes (1967) published his monograph; s<strong>in</strong>ce <strong>the</strong>n many<br />

scholars have taken up this subject. The study by Wilkes (op. cit.) is not only<br />

<strong>the</strong> best one ever made <strong>of</strong> teos<strong>in</strong>te; it also presents a taxonomy that differs from<br />

<strong>the</strong> previous one (as was seen <strong>in</strong> Chapter 1), <strong>and</strong> he separates maize as a species<br />

different from teos<strong>in</strong>te, whereas Iltis <strong>and</strong> Doebley considered it at <strong>the</strong> subspecie<br />

level, for <strong>the</strong>y based <strong>the</strong>ir work only on <strong>the</strong> compatibility <strong>of</strong> crosses <strong>and</strong> not on<br />

visible characters (Grobman, 2004: 430).<br />

Teos<strong>in</strong>te is <strong>the</strong> closest relative <strong>of</strong> maize. It has <strong>the</strong> same number <strong>of</strong> chromosomes<br />

(20), <strong>and</strong> <strong>the</strong>se are similar to those <strong>of</strong> maize <strong>in</strong> <strong>the</strong>ir lengths <strong>and</strong> <strong>in</strong><br />

<strong>the</strong> position <strong>of</strong> <strong>the</strong>ir centromeres. Teos<strong>in</strong>te hybridizes easily with maize, <strong>and</strong><br />

first-generation hybrids are vigorous <strong>and</strong> highly fertile when self-poll<strong>in</strong>ated or<br />

when <strong>the</strong>y are crossed back to ei<strong>the</strong>r parent (Mangelsdorf, 1974: 15).<br />

The first Lat<strong>in</strong> name given to teos<strong>in</strong>te was that provided by Schräder (1833),<br />

which only referred to <strong>the</strong> annual form – Euchlaena mexicana Schräder. In 1910<br />

A. S. Hitchcock (1922) discovered <strong>the</strong> perennial form <strong>of</strong> teos<strong>in</strong>te, which he called<br />

Euchlaena perennis Hitchcock. Kuntze (1904) <strong>and</strong> Reeves <strong>and</strong> Mangelsdorf<br />

(1942) subsequently ascribed it to <strong>the</strong> Zea genus <strong>and</strong> renamed it Z. mexicana<br />

(Schräder) Kuntze <strong>and</strong> Z. perennis (Hitchcock) Reeves <strong>and</strong> Mangelsdorf. 1<br />

R<strong>and</strong>olph (1976), however, disagrees with teos<strong>in</strong>te be<strong>in</strong>g ascribed to <strong>the</strong> Zea<br />

genus. He presented three tables (R<strong>and</strong>olph, op. cit.: 1 [324], 2 [325], <strong>and</strong> 3<br />

[326]) with 23 major differences <strong>in</strong> plant characteristics as well as <strong>in</strong> environmental<br />

responses between modern Zea <strong>and</strong> Euchlaena, which have enough taxonomical<br />

import to dist<strong>in</strong>guish <strong>the</strong> two genera. In a previous study (R<strong>and</strong>olph,<br />

1972; 2 see R<strong>and</strong>olph, 1976: 325), R<strong>and</strong>olph posited that <strong>the</strong>se are two different<br />

genera. In tables IV <strong>and</strong> V, R<strong>and</strong>olph (1976: 330 <strong>and</strong> 331) shows 19 differences<br />

between ancient maize <strong>and</strong> teos<strong>in</strong>te. If we ga<strong>the</strong>r <strong>the</strong> results obta<strong>in</strong>ed <strong>in</strong> <strong>the</strong><br />

analysis <strong>of</strong> both modern <strong>and</strong> archaeological specimens, <strong>the</strong>re are 32 <strong>in</strong>heritable<br />

differences between maize <strong>and</strong> teos<strong>in</strong>te.<br />

R<strong>and</strong>olph acknowledges <strong>the</strong> fact that more studies <strong>and</strong> experiments are<br />

required, but he goes aga<strong>in</strong>st <strong>the</strong> hypo<strong>the</strong>sis that teos<strong>in</strong>te is <strong>the</strong> progenitor <strong>of</strong><br />

maize (R<strong>and</strong>olph, 1976: 330). In this study R<strong>and</strong>olph presents a long list based<br />

on characteristics <strong>of</strong> genetic <strong>and</strong> taxonomic significance, which can be applied<br />

to both modern as well as archaeological maize, <strong>and</strong> which are <strong>in</strong> contrast with<br />

<strong>the</strong> essential differences <strong>of</strong> both modern teos<strong>in</strong>te uncontam<strong>in</strong>ated by maize <strong>and</strong><br />

1<br />

2<br />

For more data regard<strong>in</strong>g this po<strong>in</strong>t <strong>and</strong> full bibliographical data, see Mangelsdorf (1974:<br />

19–20) <strong>and</strong> Doebley (1990: esp. p. 7).<br />

I was unable to f<strong>in</strong>d this.

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