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a guide to the recognition and understanding of ramularia and other ...

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44<br />

PHYSIOLOGICAL LEAF SPOTS<br />

Physiological leaf spots in response <strong>to</strong><br />

disease attack<br />

For several years, plant breeders have used effective resistance genes<br />

known as mlo genes <strong>to</strong> breed varieties with resistance <strong>to</strong> powdery<br />

mildew. Some <strong>of</strong> <strong>the</strong>se varieties will induce a hypersensitive reaction<br />

when powdery mildew spores attack <strong>the</strong> plant leading <strong>to</strong> distinct leaf<br />

spots commonly known as target spots.<br />

Distinctive leaf spots associated with mildew infection on mildew resistant varieties

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