insecta mundi - Center for Systematic Entomology

14 • INSECTA MUNDI 0129, July 2010 EDMONDS AND ZIDEK
Besides the characters mentioned in the key, there are differences among the parameres of these
species that were first pointed out by Lane and Carmargo-Andrade (1935). These are not treated in the
species diagnoses below but are illustrated.
Key to species of Megaphanaeus
1. Elytral striae (Fig. 16) narrow, smooth, sides straight, converging apically. Interstriae 1-4 smooth,
devoid of coarse sculpturing (x5); interstriae 5-7 microrugose (x25), more strongly so laterally.
Female cephalic process (Fig. 27) tridentate, middle tubercle strongly elevated. Atlantic coast
forests of Brazil). bellicosus species group ............................................................................
..................................................... Coprophanaeus (Megaphanaeus) bellicosus (Olivier)
— Elytral striae widened, margins (especially of striae 2-4) undulate to some degree. Interstriae
coarsely sculptured (x5). Female cephalic process a cylindrical horn (similar to that of male)
lancifer species group ........................................................................................................... 2
2(1). Elytral interstriae (Fig. 13) at most only indistinctly granulate, usually granulorugose;
granulation, if distinct, confined to apical one-third of interstriae 2-6. Basal two-thirds of elytral
striae 2-4 moderately undulate, width at widest points about twice that of narrowest points;
strial margins at narrow points often bearing low, rounded tubercle. Metasternal shield (Fig.
18) smooth to weakly punctate; triangular, anteromedian area of metasternum (Fig. 17, arrow)
(usually hidden by retracted posterior margin of pronotum) smooth and shiny medially. Color
usually dark metallic blue. Widely distributed in Amazonia (Fig. 28) .......................................
.......................................................... Coprophanaeus (Megaphanaeus) lancifer (Linné)
— Elytral interstriae completely covered by large, round granules. Striae variable. Metasternal
shield (Fig. 19) completely punctatorugose; triangular anteromedian portion of metasternum
(usually hidden by retracted posterior margin of pronotum) finely granulate, at least laterally.
Color usually green, yellowish green, rarely bluish green. Distribution variable, but never
Amazonian ................................................................................................................................ 3
3(2). Margins of elytral striae 2-4 only gently undulate (Fig.15), strial width at narrowings at least
three-fourths of that at widest points; carinulae simple, seldom tuberculate. Triangular
anteromedian portion of metasternum finely granulate laterally, smooth and shiny medially.
Female lacking protarsi. Smaller beetles, length 25-35 mm. Xeric forests of eastern Bolivia,
Paraguay and Argentina (Fig. 28) ..............................................................................................
..................................................... Coprophanaeus (Megaphanaeus) bonariensis (Gory)
— Margins of elytral striae (Fig. 14) strongly undulate, strial width at narrowest points no more
than one-half (often much less than one-half) of that at widest points; narrowings produced by
apically tuberculate, finger-like projections of margins. Triangular anteromedian portion of
metasternum completely finely granulate, or nearly so. Female with protarsi. Large beetles,
length 30-55 mm. Dry forests of central Brazil, Bolivia, Paraguay and Argentina (Fig. 28) ....
......................................................... Coprophanaeus (Megaphanaeus) ensifer (Germar)
Coprophanaeus (Megaphanaeus) bellicosus (Olivier, 1789)
Fig. 16, 27-34
Scarabaeus bellicosus Olivier, 1789: 109
Phanaeus sylvanus Castelnau, 1840: 80 (syn. by Nevinson 1892: 2)
Megaphanaeus bellicosus (Olivier) (recomb. by Blackwelder 1944: 209)
Coprophanaeus bellicosus (Olivier) (recomb. by Edmonds 1972: 841)