42 • INSECTA MUNDI 0129, July 2010 EDMONDS AND ZIDEK Figure 113. Approximate geographic distribution of Coprophanaeus (s. str.) species in the jasius species group.
REVIEW OF COPROPHANAEUS INSECTA MUNDI 0129, July 2010 • 43 rugose. Basal pronotal fossae distinct. Elytral striae weakly, if at all, carinulate (Fig. 100). Parameres longer, length clearly greater than one-half that of phallobase (Fig. 123). Body length usually greater than 25 mm ................................. Coprophanaeus (C.) acrisius (MacLeay) 3(1). Pronotal carina of large male (Fig. 95) with central tubercle only, lacking lateral tubercles. Pronotal process of female (Fig. 125) a simple transverse carina located high on disk and flanked by shallow concavities descending toward eyes; surface sharply declivitous in front of carina. Pronotum with finely rugose, elongate depression extending from middle of posterior margin to middle of disk (Fig. 95); surface slightly swollen and smooth to finely punctate posterolaterally and along sides of medial depression. Parameres short, length barely approaching one-half that of phallobase (Fig. 128). Completely black or brown, no hint of metallic color anywhere on body surface. Atlantic coast <strong>for</strong>ests of Brazil ................. Coprophanaeus (C.) cerberus (Harold) — Pronotal carina of large male (Fig. 96, 98) clearly trituberculate (middle tubercle may be very large compared to lateral ones). Pronotal process of large female trapezoidal or quadrate (Fig. 105-106), summit usually distinctly trituberculate, set very near anterior pronotal margin and followed by oval concavity. Pronotal sculpturing variable. Color dark, often appearing black to unaided eye, but magnification (x5) under bright light always reveals metallic color (most commonly on pronotum and pygidium). Parameres longer, length clearly exceeding one-half that of phallobase ...................................................................................................................... 4 4(3). Median area of posterior portion of pronotum (Fig. 96) more-or-less smooth, any obvious sculpturing effaced; smooth area often extends anteriorly to middle of disk and laterally some distance along posterior pronotal margin. Basal pronotal fossae with sparse punctures or microgranules (x25); adjacent area sparsely, weakly punctate.................................................................................. 5 — Posteromedian area of pronotum (Fig. 98) punctate to punctatorugose; sculpturing usually weaker as it radiates anteriorly from area of basal fossae. Basal pronotal fossae and surrounding area densely, finely granulate (x15) ................................................................................................. 6 5(4). Male pronotal carina (Fig. 96) more-or-less straight, central tubercle only slightly larger than laterals. Base of male cephalic horn coarsely punctate (Fig. 103); maximum horn height not reaching level of pronotal carina (Fig. 131); median tubercle (viewed laterally) slightly inclined anteriorly (relative to base of horn). Female pronotal process (Fig. 107) trapezoidal, summit weakly trituberculate, width at summit less than interocular distance. Anteromedian pronotal concavity of female shallow, posterior (upper) margin simple. Sulcus in front of basal pronotal carina smooth (Fig. 109) ............................................. Coprophanaeus (C.) jasius (Olivier) — Middle tubercle of large-male pronotal process conspicuously larger than lateral ones, producing an acute projection or large, tab-like process (Fig. 137-138). Anterior surface of male cephalic horn minutely and rather sparsely punctured; median tooth elongate, reaching level of pronotal process (Fig. 137), more-or-less straight except slight posterior bend at apex. Pronotal process of large female (Fig. 105) strongly elevated, quadrate, summit strongly trituberculate, width at summit usually slightly greater than interocular distance. Anteromedian pronotal concavity of female deep, posterior (upper) margin weakly bitumose. Sulcus in front of basal pronotal carina (Fig. 110) sculptured .................................................. Coprophanaeus (C.) abas (MacLeay) 6(4). Pronotal process of large female strongly raised, quadrate, at most only weakly tridentate (Fig. 106). Middle tubercle of female cephalic carina strongly elevated, imparting almost triangular shape to carina (Fig. 99). Width of large-male pronotal carina equal to or greater than distance separating outer margins of eyes. Basal carinate margin of pronotum densely punctate to crenulate (Fig. 111) .................................................... Coprophanaeus (C.) gamezi Arnaud — Pronotal process of large female trapezoidal, summit weakly tridentate (as in Fig. 107). Middle tubercle of female cephalic carina only slightly larger than lateral tubercles, carina only moderately raised. Width of large-male pronotal carina less than distance separating outer margins of eyes. Basal carinate margin of pronotum entire, not broken up by strong puncturing ..................................................................... Coprophanaeus (C.) cyanescens (Olsoufieff)
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