insecta mundi - Center for Systematic Entomology

2 • INSECTA MUNDI 0129, July 2010 EDMONDS AND ZIDEK
Arnaud’s (2002c) review of the genus follows Edmonds’ subgeneric classification, except for the placement
of C. bellicosus (see Comments for C. bellicosus).
As used here, Megaphanaeus is exactly in the sense of Olsoufieff (1924), Blackwelder (1944) and
Edmonds (1972), a subgeneric group of four common species remarkable for their large size. In
Metallophanaeus Olsoufieff included but two species; Edmonds (1972) transferred into it a second species
group originally placed in Coprophanaeus s. str., and here we add several others distributed between the
two groups. The nominate subgenus originally embraced 21 species, many of which Blackwelder (1944)
erroneously transferred to Phanaeus. The subgenus Coprophanaeus s. str. is here enlarged and restructured
and the arrangement proposed by Edmonds (1972), which was based on too few species, is largely
abandoned. Arnaud’s (2002c) organization of this subgenus is different from that proposed here. Following
is a list of those 77 available (or potentially available) names here assigned to Coprophanaeus and their
present status [brackets enclose text page reference to name]:
abas MacLeay - valid name [p. 50]
acrisius MacLeay - valid name [p. 44]
ajax Sturm - junior subjective synonym of C. ensifer [p. 20]
alvarengai Pereira and d’Andretta - junior subjective synonym of C. pertyi [p. 33]
argentinus Martínez - junior subjective synonym of C. bonariensis [p. 19]
arrowi Olsoufieff - junior subjective synonym of C. dardanus [p. 80]
bellicosus Olivier - valid name [p. 14]
bitias Harold - junior subjective synonym of C. dardanus [p. 80]
bonariensis Gory - valid name [p. 19]
boucardi Nevinson - valid name [p. 65]
callegarii Arnaud - valid name [p. 97]
camargoi Pessôa - junior subjective synonym of C. acrisius [p. 44]
caroliae Edmonds - valid name [p. 102]
cerberus Harold - valid name [p. 47]
chabrillaci Thompson - junior subjective synonym of C. saphirinus [p. 29]
chiriquensis Olsoufieff - valid name [p. 71]
christophorowi Olsoufieff - valid name [p. 85]
conocephalus Olsoufieff - valid name [p. 69]
corythus Harold - valid name [p. 93]
costatus Olsoufieff - junior subjective synonym of C. cyanescens [p. 54]
cyanescens Olsoufieff - valid name [p. 54]
dardanus MacLeay - valid name [p. 80]
degallieri Arnaud - valid name [p. 87]
ducalis Castelnau - junior subjective synonym of C. ensifer [p. 20]
edmondsi Arnaud - junior subjective synonym of C. conocephalus [p. 69]
ensifer Germar - valid name [p. 20]
florenti Arnaud - junior subjective synonym of C. ohausi [p. 104]
gamezi Arnaud - valid name [p. 52]
gilli Arnaud - valid name [p. 73]
henryi Malý and Pokorný - junior subjective synonym of C. gilli [p. 73]
heros Castelnau - junior subjective synonym of C. lancifer [p. 16]
horus Waterhouse - valid name [p. 25]
ignecinctus Felsche - valid name [p. 89]
jasion Felsche - junior subjective synonym of C. dardanus [p. 80]
jasius Olivier - valid name [p. 48]
kohlmanni Arnaud - junior subjective synonym of C. morenoi [p. 60]
lancifer Linné - valid name [p. 16]
larseni Arnaud - uncertain status (see Comments under C. suredai) [p. 104]
lecromi Arnaud - uncertain status (see Comments under C. caroliae) [p. 102]
lichyi Arnaud - uncertain status (see Comments under C. ohausi) [p. 104]
machadoi Pereira and d’Andretta - junior subjective synonym of C. saphirinus [p. 29]