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The coconut odyssey - the bounteous possibilities of the ... - ACIAR

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Figure 2-2. Four common fruit colours, light brown, orange, yellow and pale green, found especially on Dwarf palms.<br />

Many o<strong>the</strong>r shades <strong>of</strong> green and brown are found on Tall fruit but <strong>the</strong> recessive orange and yellow colours shown here are<br />

almost exclusive to Dwarf palms, which display <strong>the</strong> same colours on <strong>the</strong> frond petiole and also have pale leaflets. (Roland<br />

Bourdeix, Centre for International Cooperation in Agricultural Research for Development (CIRAD), France)<br />

a result <strong>of</strong> selection by humans over<br />

many millennia for greater water<br />

content, providing a convenient<br />

water supply for drinking where<br />

fresh water is scarce, especially on<br />

sea voyages.<br />

�e ‘wild’ form <strong>of</strong> <strong>the</strong> Tall <strong>coconut</strong>,<br />

on <strong>the</strong> o<strong>the</strong>r hand, is found wherever<br />

<strong>the</strong> palm has spread naturally,<br />

and has remained genetically<br />

unchanged in its proportions <strong>of</strong><br />

husk and nut. �e wild-type fruit<br />

is smaller than <strong>the</strong> domesticated<br />

fruit and has a higher proportion<br />

<strong>of</strong> husk, which enables it to float<br />

on <strong>the</strong> ocean for a long time, and a<br />

correspondingly lower volume <strong>of</strong><br />

nut-water. Intermediate-sized fruits,<br />

showing great diversity between<br />

palms, are found in many <strong>coconut</strong><br />

populations. �is is especially so on<br />

some <strong>of</strong> <strong>the</strong> long-occupied Pacific<br />

islands, indicating that an original<br />

wild <strong>coconut</strong> palm population,<br />

which predated <strong>the</strong> arrival <strong>of</strong> human<br />

colonisers, has been blending for<br />

some time with introduced largerfruited<br />

varieties.<br />

A few very remote islands, such as<br />

North Keeling in <strong>the</strong> Cocos Islands<br />

<strong>of</strong> <strong>the</strong> Indian Ocean, show truly<br />

wild-type fruit with no trace <strong>of</strong><br />

introduced forms.<br />

36<br />

Drought tolerance<br />

Apart from <strong>the</strong>ir recognisable fruit<br />

characters (size, shape, dominant<br />

colour), different Tall populations<br />

can also be distinguished, when<br />

compared within a location, through<br />

<strong>the</strong>ir differences in adaptation.<br />

Environmental variables (such<br />

as <strong>the</strong> regular occurrence <strong>of</strong> a<br />

dry season, seasonal variations in<br />

temperature, or attacks by pest and<br />

disease organisms) induce tolerance<br />

through natural selection. For<br />

example, <strong>coconut</strong> palms from Kerala,<br />

in south-west India where <strong>the</strong> dry<br />

season is pronounced, withstand<br />

<strong>the</strong> seasonal drought well at home.

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