The coconut odyssey - the bounteous possibilities of the ... - ACIAR
The coconut odyssey - the bounteous possibilities of the ... - ACIAR
The coconut odyssey - the bounteous possibilities of the ... - ACIAR
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over during this concealed phase<br />
<strong>of</strong> germination, as <strong>the</strong> direction<br />
<strong>the</strong> roots and shoot would become<br />
distorted and <strong>the</strong>ir growth would be<br />
retarded. �e shoot tip requires two<br />
to four weeks to elongate sufficiently<br />
to appear through <strong>the</strong> husk,<br />
depending on its vigour and on <strong>the</strong><br />
thickness <strong>of</strong> <strong>the</strong> husk. Slicing <strong>of</strong>f a<br />
section <strong>of</strong> husk immediately above<br />
<strong>the</strong> s<strong>of</strong>t eye hastens emergence.<br />
In <strong>the</strong> purposeful management <strong>of</strong><br />
<strong>the</strong> seedling, <strong>the</strong> early phase is called<br />
<strong>the</strong> ‘pre-nursery’. It is followed by <strong>the</strong><br />
planting <strong>of</strong> <strong>the</strong> developing seedling<br />
into a large polybag, by which time<br />
<strong>the</strong> roots have probably emerged<br />
from <strong>the</strong> underside <strong>of</strong> <strong>the</strong> husk,<br />
sometimes struggling to force <strong>the</strong>ir<br />
tips through <strong>the</strong> thin, but ra<strong>the</strong>r<br />
tough, outer layer. �is layer is best<br />
sliced away to encourage rapid root<br />
emergence. If <strong>the</strong> nut is large, more<br />
<strong>of</strong> <strong>the</strong> husk might need to be sliced<br />
<strong>of</strong>f to fit it more readily into <strong>the</strong><br />
polybag, submerged to half its depth<br />
in <strong>the</strong> potting soil.<br />
�e seedling ‘sponges’<br />
from <strong>the</strong> kernel<br />
�e haustorium—<strong>the</strong> s<strong>of</strong>t, spongy<br />
internal organ expanding within<br />
<strong>the</strong> nut—is <strong>of</strong>ten referred to as<br />
<strong>the</strong> ‘egg’ or ‘apple’. Progressively, <strong>the</strong><br />
haustorium makes contact with <strong>the</strong><br />
entire inner surface <strong>of</strong> <strong>the</strong> kernel<br />
(except in very large nuts), releasing<br />
s<strong>of</strong>tening enzymes as it does so. It<br />
rapidly absorbs <strong>the</strong> nut water, and<br />
from <strong>the</strong> kernel it takes nutrients<br />
at an increasing rate to provide<br />
substance and energy for <strong>the</strong> growth<br />
<strong>of</strong> <strong>the</strong> seedling.<br />
In <strong>the</strong> three-month gap between<br />
germination and <strong>the</strong> expansion<br />
<strong>of</strong> <strong>the</strong> first green leaf, <strong>the</strong> seedling<br />
depends solely on <strong>the</strong> kernel for its<br />
energy supply. �ere follows a period<br />
<strong>of</strong> gradual ‘weaning’ as <strong>the</strong> young<br />
seedling expands a series <strong>of</strong> leaves,<br />
46<br />
each larger than <strong>the</strong> one before. Each<br />
new leaf supplies a greater share <strong>of</strong><br />
<strong>the</strong> energy needs <strong>of</strong> <strong>the</strong> seedling,<br />
which <strong>the</strong>reby becomes less and less<br />
reliant on <strong>the</strong> diminishing kernel to<br />
sustain its accelerating growth.<br />
�e rich nutrient and energy<br />
mixture derived from <strong>the</strong> kernel<br />
is delivered by <strong>the</strong> haustorium to<br />
<strong>the</strong> developing seedling for at least<br />
12 months, although sooner or<br />
later a proportion <strong>of</strong> seeds suffers<br />
invasion by fungi or bacteria that<br />
disable <strong>the</strong> haustorium. While <strong>the</strong><br />
seedling is able to survive this loss<br />
<strong>of</strong> access to <strong>the</strong> kernel, it suffers a<br />
Figure 3-4. Polybag seedlings in a large plantation nursery in Mozambique<br />
(1992). �e seedlings will be transplanted to <strong>the</strong> field without any significant<br />
setback after developing three or four more leaves.