Gliding Mammals: Taxonomy of Living and Extinct Species

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26 • SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY Wilson (1993:135) or Stafford and Szalay (2000:360). Synonymized within variegatus by Stafford (2005:110). Galeopterus variegatus temminckii (Waterhouse, 1839) [Galeopithecus] temminckii Waterhouse, 1839b:119. type Locality. Sumatra. comments. Tuangku Island (Banyak Islands) population erroneously named Galeopterus volans by Miller (1903b:471). Incorrectly synonymized within volans by Wilson (1993:135). Valid subspecies according to Chasen and Kloss (1929a:16), Chasen (1940:20), and Stafford and Szalay (2000:360). Synonymized within variegatus by Corbet and Hill (1992:53) and Stafford (2005:110). distribution. Sumatra, Lingga Islands (Bakung, Penuba, Sebangka, and Singkep islands) off eastern Sumatra, Bangka Island, and Rupat Island in the Strait of Malacca, Indonesia (Stafford and Szalay, 2000:360; Meijaard, 2003:1256). Also occurs on Tuangku Island and Bankaru Island (Banyak Islands), Musala Island, and Batu Islands (Tello, Pini, Tana Bala, and Tana Masa islands) off western Sumatra (Meijaard, 2003:1256). Galeopithecus marmoratus Temminck, 1836:ix. type Locality. Sumatra, Indonesia. comments. Synonymized within volans by Wilson (1993:135). Stafford (2005:110) notes that Cabrera (1925:210) listed “G. marmoratus Temminck (1829)” as a synonym of G. variegatus, but Stafford was unable to locate that description, and the citation is unknown. Fischer (1829:79) lists this taxon as “G.[aleopithecus] marmorati nomen ab illo obtigit.” Name and type locality are discussed by Mees (1957:215). Galeopithecus saturatus Miller, 1903a:51. type Locality. Tana Bala Island, Batu Islands, western Sumatra, Indonesia. comments. Recognized as a subspecies of variegatus by Chasen (1940:20). Synonymized within variegatus by Corbet and Hill (1992:53), Wilson (1993:135), and Stafford (2005:110). Galeopithecus tuancus Miller, 1903a:53. type Locality. Tuangku Island, Banyak (Banjak) Islands, western Sumatra, Indonesia. comments. Recognized as a subspecies of variegatus by Chasen (1940:20). Synonymized within variegatus by Corbet and Hill (1992:53), Wilson (1993:135), and Stafford (2005:110). Cynocephalus tellonis Lyon, 1908:139. type Locality. Tello Island, Batu Islands, western Sumatra, Indonesia. comments. Recognized as a subspecies of variegatus by Chasen (1940:20). Synonymized within variegatus by Corbet and Hill (1992:53), Wilson (1993:135), and Stafford (2005:110). †Dermotherium Ducrocq et al., 1992 †Dermotherium Ducrocq et al., 1992:373. type species. †Dermotherium major Ducrocq et al., 1992:373. comments. Late Eocene, Asia. Placed within the family Galeopithecidae by McKenna and Bell (1997: 328). †Dermotherium major Ducrocq et al., 1992 †Dermotherium major Ducrocq et al., 1992:373. type Locality. Wai Lek lignite pit, Changwat Krabi, southern Thailand. comments. Late Eocene. Stafford and Szalay (2000:378) suggest that if this species proves to be a dermopteran, then it may be appropriate to place this species within the family Cynocephalidae; however, they suggest the Dermopteran affinities are uncertain as the specimen is poorly preserved and only m3 has any diagnostic characters. These authors also suggest that some of the features listed as characteristic of Dermoptera are more widespread than is acknowledged by Ducrocq et al. (1992). †Dermotherium chimaera Marivaux et al., 2006 †Dermotherium chimaera Marivaux et al., 2006:398. type Locality. Cha Prong pit, Nong Ya Plong coal mine, Phetchaburi Province, northern part of the Thailand peninsula. comments. Late Oligocene. Specimens close to this species have also been discovered from early Eocene deposits in Pakistan (Marivaux et al., 2006). †faMily plagioMenidae Matthew, 1918 †Family Plagiomenidae Matthew, 1918:598. type genus. †Plagiomene Matthew, 1918: 601. comments. Early Paleocene to mid- Eocene, late? Eocene, late Oligocene, North America. Family placed within Dermoptera by McKenna and Bell (1997:327) and
Silcox et al. (2005:134), with further support added by Bloch et al. (2007:1163). †Elpidophorus Simpson, 1927 †Elpidophorus Simpson, 1927:5. type species. †Elpidophorus elegans Simpson, 1927:5. comments. Early to late Paleocene, North America. Genus was tentatively placed within the †family Oxyclaenidae by Simpson (1927:2), within Dermoptera by Gunnell (1989:28), and within †Plagiomenidae by Rose (1975:676), McKenna and Bell (1997:327), and Bloch et al. (2007:1163). †Elpidophorus elegans Simpson, 1927 †Elpidophorus elegans Simpson, 1927:5. type Locality. Paskapoo Formation, Alberta, Canada. comments. Early to late Paleocene. †Elpidophorus patratus Simpson, 1936:11. type Locality. Scarritt Quarry, Fort Union, Crazy Mountain Field, Montana, USA. comments. Upper Paleocene. Synonymized within †elegans by Szalay (1969:218, 219) and Rose (1975:676). †Elpidophorus minor Simpson, 1937 †Elpidophorus minor Simpson, 1937:133. type Locality. Probably Silberling Quarry, Fort Union, Crazy Mountain Field, Montana, USA. comments. Middle Paleocene. †Eudaemonema Simpson, 1935 †Eudaemonema Simpson, 1935:231. type species. †Eudaemonema cuspidata Simpson, 1935:231. comments. Middle Paleocene. North America. Genus placed within the †family Mixodectidae by Gunnell (1989:56) and within the †family Plagiomenidae by McKenna and Bell (1997:327). †Eudaemonema cuspidata Simpson, 1935 †Eudaemonema cuspidata Simpson, 1935:231. type Locality. Fort Union, Montana, USA. comments. Middle Paleocene. NUMBER 638 • 27 †subfaMily plagioMeninae Matthew, 1918 †Family Plagiomenidae Matthew, 1918:598. type genus. †Plagiomene Matthew, 1918:601. comments. Late Paleocene to early Eocene, North America. Subfamily rank recognized by Russell et al. (1973:50), McKenna (1990:212), and McKenna and Bell (1997:327). Included within Dermoptera by McKenna and Bell (1997:327) and Sundatheria by Bloch et al. (2007:1162). †tribe worlandiini bown and rose, 1979 †Tribe Worlandinae Bown and Rose, 1979:97. type genus. †Worlandia Bown and Rose, 1979:97. comments. Late Paleocene to early Eocene, North America. Recognized at tribe rank by McKenna (1990:212, 231) and McKenna and Bell (1997:327). †Planetetherium Simpson, 1928 †Planetetherium Simpson, 1928:11. type species. †Planetetherium mirabile Simpson, 1928:11. comments. Late Paleocene, North America. Genus recognized by Rose and Simons (1977:221) and Bown and Rose (1979:89). †Planetetherium mirabile Simpson, 1928 †Planetetherium mirabile Simpson, 1928:11. type Locality. Fort Union, southern Montana, USA. comments. Late Paleocene, Park County, Wyoming, USA. †Worlandia Bown and Rose, 1979 †Worlandia Bown and Rose, 1979:97. type species. †Worlandia inusitata Bown and Rose, 1979:99. comments. Late Paleocene to early Eocene, North America. Genus placed within Dermoptera by Gunnell (1989:147) and the †family Plagiomenidae by Bown and Rose (1979:97), Gingerich (1987:311), and McKenna and Bell (1997:327). †Worlandia inusitata Bown and Rose, 1979 †Worlandia inusitata Bown and Rose, 1979:99. type Locality. Park County, Wyoming, USA. comments. Early Eocene.
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