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Introduction to Cladistic Analysis - Integrative Biology - University of ...

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Copyright © 2009 by Department <strong>of</strong> <strong>Integrative</strong> <strong>Biology</strong>, <strong>University</strong> <strong>of</strong> California-Berkeley<br />

unnamed, but any branch that connects two nodes (remember, a node represents a<br />

speciation event) corresponds <strong>to</strong> an ances<strong>to</strong>r (technically, an internodal branch may<br />

correspond <strong>to</strong> a series <strong>of</strong> ances<strong>to</strong>rs).<br />

Figure B is a very small cladogram. The following cladogram (Figure C) was constructed<br />

for the six taxa from Figure A:<br />

Figure C. A cladogram.<br />

tunicate lamprey Cladoselache † trout lungfish frog<br />

1<br />

i<br />

2<br />

Besides the six taxa <strong>of</strong> interest, this cladogram displays five speciation events (labeled i-v)<br />

and five hypothetical ances<strong>to</strong>rs (labeled 1-5).<br />

Ances<strong>to</strong>r 5 is the most recent common ances<strong>to</strong>r <strong>of</strong> lungfish and frog.<br />

Lungfish and frog are more closely related <strong>to</strong> each other than either is <strong>to</strong> any<br />

<strong>of</strong> the other four taxa under study. Ances<strong>to</strong>r 5, lungfish, and frog <strong>to</strong>gether<br />

compose a clade (monophyletic group).<br />

Similarly, Ances<strong>to</strong>r 4 is the most recent common ances<strong>to</strong>r <strong>of</strong> trout, lungfish,<br />

and frog. Trout, lungfish, and frog are more closely related <strong>to</strong> each other than<br />

either is <strong>to</strong> any <strong>of</strong> the other three taxa included in the analysis. Ances<strong>to</strong>r 4,<br />

trout, lungfish, and frog <strong>to</strong>gether compose a second clade.<br />

This cladogram contains three more clades (monophyletic groups). Can you identify them?<br />

<strong>Introduction</strong> <strong>to</strong> <strong>Cladistic</strong> <strong>Analysis</strong> - 4<br />

ii<br />

3<br />

iii<br />

Cladograms and time<br />

A cladogram provides information about the relative timing <strong>of</strong> speciation events (and, thus,<br />

the relative antiquity <strong>of</strong> ances<strong>to</strong>rs). Look again at the cladogram shown in Figure C.<br />

4<br />

iv<br />

5<br />

v

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