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AMAZONAS<br />

28<br />

base of the dorsal fin. These spots are almost always surrounded by bright blue. This angel<br />

always has a red eye and the mouth is turned slightly upward. There are six or seven stripes<br />

visible on the dorsal fin, while those on the caudal fin are only rarely apparent.<br />

Pterophyllum sp. 4: To date I have caught this form only once, and that was in the Río<br />

Demini drainage (a tributary of the Río Negro). It is the only angel I have found so far<br />

that exhibits eight striking bands.<br />

Pterophyllum sp. 5: This angel, which I found in the ichthyologically unexplored Río<br />

Apaporis (Colombia), is distinguished from all other angelfishes by its small number of<br />

dorsal-fin rays. Other differences include its large, silvery scales, the striking, large humeral<br />

spot immediately behind the eye, and the very irregular eight or nine stripes, often<br />

more like spots, on the dorsal fin. By contrast, the caudal fin has only three to four broad<br />

reddish stripes.<br />

Pterophyllum sp. 6: Like Pterophyllum sp. 5, the angel that I found in the Lago Aipauá<br />

(Río Purus basin) has larger scales. But a very striking feature is the extremely wide<br />

posterior black band extending from the extreme end of the dorsal<br />

to the last ray of the anal fin. In addition, there are five to six<br />

broad black stripes clearly visible on the dorsal, somewhat<br />

less striking on the caudal fin. The pectoral fins are the<br />

longest I have ever seen in any angelfish except Pterophyllum<br />

altum.<br />

Pterophyllum altum: This species was<br />

described by Pellegrin in 1903. Natasha<br />

Khardina and I have examined the<br />

specimens in the Paris Museum (the jar<br />

apparently hadn’t been opened since 1903).<br />

The three specimens originated from the lower<br />

Atabapo, Río Orinoco, Venezuela.<br />

Pterophyllum scalare: This species was described<br />

by Schultze in 1823 on the basis of a single specimen<br />

collected by M.H.C. Lichtenstein, zoologist and<br />

first director of the Berlin Zoo. Here I illustrate for the first<br />

time the variants that I assign to this species—all with precise<br />

collection site data. This species is often labeled as Pterophyllum<br />

altum (Río Negro Altum) in the aquarium hobby and sometimes in the<br />

Pterophyllum scalare from the Río Negro in Brazil.<br />

Pterophyllum scalare<br />

from French Guiana,<br />

possibly a distinct<br />

species?<br />

TOP: AQUAPRESS/PLANQUETTE; BOTTOM: H. BLEHER; OPPOSITE PAGE, BOTTOM RIGHT: N. KHARDINA; OTHERS: H. BLEHER

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