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AMAZONAS<br />

50<br />

Despite this preservation shortfall in domesticated angelfishes,<br />

one thing has become clear: in a world where<br />

the community ethic has embraced the open-sourcing of<br />

genetic knowledge, any angelfish breeder with the genetic<br />

raw materials can recreate and refine the desired genetic<br />

result.<br />

Angelfish genetics:<br />

a history of standardization<br />

In no small part, the late Dr. Joanne Norton is credited<br />

with laying the foundation for most of the current<br />

understanding of angelfish genetics. From 1982 to 1994,<br />

Norton published no fewer than 18 articles in Freshwater<br />

and Marine Aquarium (FAMA) covering her extensive<br />

personal work divining the genetic basis for the myriad<br />

of forms being produced around the world. It is fair<br />

to say that Dr. Norton demonstrated how an aquarist<br />

can single-handedly change the aquarium hobby and<br />

industry, and, without a doubt, the tradition of genetic<br />

transparency was indoctrinated through Dr. Norton’s<br />

visionary contributions.<br />

The Angelfish Society (www.theangelfishsociety.org,<br />

TAS) was founded in 2000 and incorporated in 2003 as<br />

Classic Forms<br />

Top row, left to right:<br />

Halfblack Veil Angelfish (V/+<br />

- h/h), Black Angelfish (D/g)<br />

(a.k.a. Hybrid Black),<br />

Koi Veil Angelfish<br />

(Gm/g - S/S - V/+)<br />

Philippine Blue Influences<br />

Bottom row, left to right:<br />

Pinoy Veil WiFi (Widefin)<br />

Ghost Angelfish (D/+ - S/+<br />

- V/+ - pb/pb), Blue Marble<br />

Angelfish (M/+ - pb/pb)<br />

a not-for-profit organization. While the Society provides<br />

and performs many functions, it also perpetuates the<br />

tradition of Dr. Norton’s work. The current website includes<br />

a repository of the 18 seminal works she published<br />

in FAMA, but takes it a step further with the phenotypes<br />

library that’s readily available online (http://www.theangelfishsociety.org/phenotype_library_2007/NewIndex.<br />

html). This phenotype library covers all the known genetics<br />

as uncovered by Dr. Norton in the 1980s and ’90s,<br />

and was last fully updated in 2007.<br />

This genetic canon is generally well regarded as a<br />

definitive starting point, although observable traits may<br />

be the result of multiple genes or what breeder Frank<br />

O’Neill routinely refers to as “genetic modifiers.” Then<br />

there are occasional mysteries that stand in stark contrast<br />

to the assumed-proven genetics. Current TAS president<br />

Tamar Stephens points out, “The genes that we have<br />

listed [in our phenotype library] are all genes that follow<br />

Mendelian genetics. With multigenic traits, multiple<br />

genes contribute to the overall effect, making the results<br />

unpredictable.” That unpredictability may be the reason<br />

why many traits have yet to reveal their genetic secrets.<br />

The Angelfish Society has taken on the role of the<br />

MATT PEDERSEN; BLACK ANGELFISH: MELLOW AQUATICS

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