Bryological Monograph An annotated checklist of the mosses of ...

ANNOTATIONS
1. Ignatov & Afonina (1992) report Sphagnum imbricatum
Hornsch. ex Russow (section Sphagnum), S.
orientale L.I.Savicz (section Subsecunda) and S.
perfoliatum L.I.Savicz (section Subsecunda) from
arctic European Russia. The record of S. imbricatum
would, if correct, be likely to refer to S. steerei
R.E.Andrus, which is seemingly the only taxon of the
S. imbricatum complex collected in arctic areas of
North America and Eurasia (Flatberg, 1984), and
which is listed by Dierssen (2001). However, Afonina
(pers. comm. to Flatberg) tells us that there are no
reliable herbarium specimens of European material of
any of these species. They are therefore omitted.
2. Sphagnum viride is closely related to S. cuspidatum
(Flatberg, 1988), and isoenzymatic results indicate
that gene flow exists between the two morphs
(Hanssen, Sa˚stad & Flatberg, 2000). More genetic
data are required to evaluate their taxonomic status.
3. Sphagnum brevifolium and S. isoviitae belong to a
small group of closely related taxa in the S. recurvum
complex with yellow spores (Flatberg, 1992, 1992
[1993]). In typical morphological appearance they are
well separated from S. fallax, but seem not clearly
segregated genetically in sympatric populations
(Sa˚stad, Stenøien & Flatberg, 1999). Their species
status can therefore be questioned, and more genetic
data are required before a decisive taxonomic conclusion
can be drawn.
4. The name Sphagnum lenense was published as a
nomen nudum in 1915, and was not validated until
1936 (Afonina and Egorova, pers. comm. to Flatberg).
5. Sphagnum auriculatum is retained here as the name for
the taxon that has, following a paper by Dirkse &
Isoviita (1986), been called S. denticulatum in recent
lists. Sphagnum denticulatum is an aquatic form with
abnormally large, nearly isophyllous leaves. Although
most such plants belong to S. auriculatum, similar
forms of S. inundatum are found not uncommonly
growing under aquatic conditions in mire pools along
the western coast of Norway. The type specimen of S.
denticulatum is therefore unsatisfactory.
6. Sphagnum tundrae was described from Svalbard by
Flatberg (1994).
7. Sphagnum wulfianum is either nested within section
Acutifolia (Shaw, 2000b) or is sister to section
Acutifolia, lying between it and section Squarrosa
(Shaw, Cox & Boles, 2005). It is left here in its own
section Polyclada.
8. Sphagnum aongstroemii may be nested within section
Acutifolia (Shaw, 2000b, Shaw et al., 2005). It is left
here in its own section Insulosa.
9. Sphagnum nitidulum Warnst. was described from
Terceira in the Azores, but type material has
presumably been destroyed and its identity is obscure.
MOSSES OF EUROPE AND MACARONESIA 227
10. Sphagnum subtile (Russow) Warnst. (S. capillifolium
var. subtile (Russow) Kartt., S. rubellum var. subtile
(Russow) Amann) is a disputed taxon whose distinctness
can be questioned. Natcheva & Cronberg (2002)
consider European herbarium material studied to
belong to S. rubellum. Shaw et al. (2005) claim that
North American plants named S. subtile cannot be
separated genetically from S. capillifolium. The name
is based on European material, but original material
has not been traced. It is premature to disregard the
existence of a European taxon underlying this name.
11. Sphagnum tenerum Sull. & Lesq. ex Sull. (S. capillifolium
var. tenerum (Sull. & Lesq. ex Sull.) H.A.Crum)
is an eastern North American species, which according
to Shaw et al. (2005) is highly differentiated genetically
from S. capillifolium. European plants labelled S.
tenerum have been examined by Flatberg, and belong
to hemi-isophyllous forms of S. capillifolium.
12. Sphagnum olafii was described from Svalbard by
Flatberg (1993a).
13. Sphagnum rubiginosum was described from Norway by
Flatberg (1993b).
14. Sphagnum andersonianum has been reported from
Europe, but Shaw et al. (2005) claim that examined
material from Europe and North America cannot be
distinguished by genetic markers from S. rubellum.
15. Sections of Andreaea follow Murray (1988).
16. Sérgio (2004) recognized Andreaea heinemannii subsp.
crassifolia, stating that it does not intergrade morphologically
with subsp. heinemannii and that they are
found in one mixed population. The taxon may thus
merit species status.
17. Smith (2004) treated the East Asian Atrichum rhystophyllum
(Müll.Hal.) Paris as a synonym of A.
angustatum, following Richards & Wallace (1950),
who treated it as a variety of A. angustatum. The
relationships of North American, European and East
Asian representatives of this complex and their
affinity with A. undulatum are not clear and are in
need of thorough revision, as stated by Lou &
Koponen (1986).
18. A critical, global revision is required to clarify the
status of Atrichum flavisetum and its relationship to A.
undulatum.
19. Pogonatum inflexum (Lindb.) Sande Lac. is listed for
Turkey by Kürschner & Erdag˘ (2005). Pogonatum
inflexum is an exclusively East Asian species confined
to China, Japan, Korea and the Russian Far East.
Several records from other parts of mainland Asia
have been confirmed as misidentified, and are mostly
P. neesii (Hyvönen, 1989). We have not studied any
Turkish material but are inclined to assume that these
records also represent P. neesii, a widespread Asian
species with a disjunct occurrence in the Caucasus
area (see below).
20. According to Schratz (1928), Pogonatum aloides var.
minimum (Crome) Molendo and P. nanum var. long-