Bryological Monograph An annotated checklist of the mosses of ...

393. Hypnum holmenii was described from Canada,
Greenland and Finland (Ando, 1994), and has been
found in the Arkhangelsk Oblast of Russia (Ignatov &
Ignatova, 2004).
394. Hypnum subimponens is reported from European
Russia by Afonina (2004).
395. Vesicularia sphaerocarpa (A.Jaeger) Broth. was
reported from Malta by Reimers in 1934 on the basis
of a 19th century collection by Schweinfurth. The
specimen (B) was destroyed in the Second World War,
but Reimers in Bizot & Potier de la Varde (1952) was
confident that it belonged to the taxon described by
these authors as V. reimersiana. According to Corley
et al. (1981), V. sphaerocarpa is a synonym of V.
galerulata (Duby) Broth. Düll (1985) listed the
Maltese plant as V. reimersiana, but also asserted that
V. galerulata had been found in mainland Italy.
Corley & Crundwell (1991) accordingly added V.
reimersiana to the list, retaining V. galerulata.
However, no species of Vesicularia is listed for Italy
by Cortini Pedrotti (2006 [2005]). It seems likely that
V. galerulata has never been found in Europe.
396. Smith (2004) treated Heterocladium flaccidum as a
species, but provided only a nomen nudum. He has
made the necessary new combination above for this
checklist, noting (pers. comm. to Hill) that consultation
with a number of British field bryologists revealed
that, with one exception, none had encountered
intermediates between it and H. heteropterum.
397. Heterocladium wulfsbergii is discussed in detail by
Crundwell & Smith (2000).
398. Iwatsukiella leucotricha was listed for Europe by
Podpěra (1954) under the name Habrodon leucotrichus
(Mitt.) Perss., with a locality in the southern Urals. It
was omitted without comment by Corley et al.
(1981), but is indicated from the European side of
the Urals by Dierssen (2001) and Ignatov & Ignatova
(2004).
399. Alpine and arctic forms of Hylocomium splendens have
been variously treated. Var. alpinum Schlieph. ex
Limpr. is mapped by Söderström (1996), but is
doubted and treated as possibly a synonym of var.
obtusifolium (Geh.) Paris by Koperski et al. (2000).
Var. obtusifolium is commonly treated as a synonym
of H. alaskanum (Lesq. & James) Austin, whose
distinguishing characters have been shown by Ross et
al. (2001) to be mainly under environmental rather
than genetic control. It is not clear whether any of
these forms is sufficiently distinct genetically to
deserve taxonomic recognition.
400. Herzogiella adscendens (Lindb.) Z.Iwats. & W.B.Schofield
has been reported from Svalbard (Corley &
Crundwell, 1991; Düll, 1985), but the record
was subsequently rejected (Frisvoll & Elvebakk,
1996).
401. Myurella julacea var. ciliata (Chal.) Ochyra &
Bednarek-Ochyra (syn. M. julacea var. scabrifolia
MOSSES OF EUROPE AND MACARONESIA 243
Lindb. ex Limpr.) is treated as distinct in some
European countries, but intergrades completely with
var. julacea. It is not recognized here.
402. Orthothecium chryseon is incorrectly listed as O.
chryseum by Crosby et al. (1999). It was described as
Hypnum chryseon by Schwägrichen (1804).
403. Plagiothecium curvifolium is sometimes treated as a
variety (in Europe) or synonym (in North America)
(Ireland, 2003) of P. laetum, to which it is undoubtedly
very close. As a variety, its correct name is P. laetum
var. secundum.
404. Plagiothecium denticulatum var. undulatum is treated
in many European lists as a species, P. ruthei, but in
North America merely as a synonym of P. denticulatum
(Ireland, 2003). It is not clearly distinct from var.
denticulatum, and we follow Ignatov & Ignatova
(2004), Koperski et al. (2000) and Smith (2004) in
treating it as a variety.
405. Plagiothecium noricum was described from the
Pinzgau, Austria and was thought by Molendo
(but not Limpricht) to intergrade with P. neckeroideum.
Grims (1999) listed it but noted that Düll
(1992) had thought it a weak species, possibly
only a variety of P. neckeroideum. It is not recognized
here.
406. Plagiothecium succulentum is a difficult taxon,
intergrading on the one hand with P. nemorale
(Hemeric, 1989) and on the other with P. cavifolium
(Nyholm, 1965). It is also very close to P. platyphyllum.
It is retained here, because it cannot satisfactorily
be subordinated to any of these species without
treating the whole group as a single, rather-broad
species.
407. Plagiothecium svalbardense was described by Frisvoll
in Frisvoll & Elvebakk (1996).
408. Pseudotaxiphyllum laetevirens was raised to species
rank by Hedenäs (1992a).
409. Entodon challengeri occurs in European Russia
(Ignatov & Ignatova, 2004) and is the correct name
for the plant formerly known as E. compressus
(Iwatsuki & Tan, 2001).
410. According to Ochyra & Ireland (2004), Isopterygium
tenerum is absent from Europe but present in sub-
Saharan Africa. However, Schumacker in Stewart
(1995), Cortini Pedrotti (2006 [2005]) and Gabriel
et al. (2005) list it for Italy and the Azores.
Isopterygium bottinii was excluded from Sematophyllum
by Guerra & Gallego (2005), and is included
here in I. tenerum.
411. Hageniella micans, previously known from Europe
and North America, was synonymized with H.
pacifica Broth., which occurs in China and Taiwan
(Tan & Jia, 1999).
412. Sematophyllum adnatum was found in Italy, where it is
thought to be an introduction from North America
(Brusa in Blockeel et al., 2000).