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Bryological Monograph An annotated checklist of the mosses of ...

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139. Molecular phylogenies <strong>of</strong> Pottiaceae by Werner et al.<br />

(2004a, 2005b) place <strong>the</strong> European species in <strong>the</strong> three<br />

groups recognized here: Merceyoideae (Scopelophila),<br />

Trichostomoideae (including Splachnobryum,<br />

Ephemeraceae, Pleuroweiseae sensu Corley et al.,<br />

1981, and some Barbula species, e.g. B. bolleana),<br />

and Pottioideae (including Pottieae, Cinclidotus and<br />

Barbuleae sensu Corley et al., 1981). Timmiella does<br />

not belong in Pottiaceae but is listed here until a better<br />

place can be found for it. Likewise, those Barbula<br />

species that belong in Trichostomoideae have been<br />

retained in Barbula until <strong>the</strong>ir correct generic position<br />

can be ascertained.<br />

140. Crumia latifolia (Kindb.) W.B.Sch<strong>of</strong>ield is known<br />

from <strong>the</strong> Caucasus but outside Europe.<br />

141. <strong>An</strong>oectangium handelii is known from <strong>the</strong> Crimea. It is<br />

described by Zander & Weber (2005), correcting an<br />

earlier report by Zander (1977) who had treated it as a<br />

synonym <strong>of</strong> Molendoa sendtneriana.<br />

142. Ephemerum hibernicum was described from Ireland by<br />

Holyoak & Bryan (2005).<br />

143. Holyoak has examined much material <strong>of</strong> Ephemerum<br />

serratum and E. minutissimum, and finds that in both<br />

species occasional plants have leaves with a weak<br />

nerve. He has not seen <strong>the</strong> type <strong>of</strong> E. serratum var.<br />

praecox, which could be a synonym <strong>of</strong> ei<strong>the</strong>r <strong>of</strong> <strong>the</strong>se<br />

taxa.<br />

144. Ephemerum spinulosum has been found in Ireland and<br />

<strong>the</strong> Basque region <strong>of</strong> Spain (Holyoak, 2001; Infante &<br />

Heras, 2005).<br />

145. Author citation for Eucladium verticillatum follows <strong>the</strong><br />

typification by Ochyra & Zijlstra (2005).<br />

146. Gymnostomum mosis (Lorentz) Jur. & Milde was<br />

reported from Spain by Martínez-Sánchez, Ros &<br />

Guerra (1991), but later Cano, Ros & Guerra (1994)<br />

described <strong>the</strong> new species Gymnostomum lanceolatum<br />

based on this material.<br />

147. Gymnostomum aeruginosum var. obscurum was<br />

described by Guerra (2004).<br />

148. Spelling <strong>of</strong> species epi<strong>the</strong>t in Hymenostylium recurvirostrum<br />

follows Hedwig (1801) not Crosby et al.<br />

(1999).<br />

149. Pleurochaete malacophylla (Müll.Hal.) Broth. is listed<br />

for Turkey by Kürschner & Erdag˘ (2005).<br />

150. Splachnobryum delicatulum is treated by Arts (2001) as<br />

a synonym <strong>of</strong> S. obtusum. Splachnobryum obtusum was<br />

not included in <strong>the</strong> main list <strong>of</strong> Corley et al. (1981),<br />

because <strong>the</strong>se authors considered that its occurrences<br />

were not sufficiently natural. However, it occurs in <strong>the</strong><br />

open in Hungary and <strong>the</strong> Azores, and is accordingly<br />

listed here.<br />

151. Tortella arctica (Arnold) Crundw. & Nyholm has been<br />

reported from Svalbard, but <strong>the</strong> records refer to<br />

Trichostomum arcticum (Frisvoll & Elvebakk,1996).<br />

152. Tortella alpicola was reported new for Europe by<br />

Otnyukova et al. (2004).<br />

MOSSES OF EUROPE AND MACARONESIA 233<br />

153. Tortella bambergeri was not recognized as a good<br />

species by Corley et al. (1981), but is generally<br />

accepted by bryologists in central Europe and also<br />

by us.<br />

154. Tortella densa is treated as a variety T. inclinata var.<br />

densa, following Eckel (1998).<br />

155. Tortella limbata has been refound on <strong>the</strong> Canary<br />

Islands. Dirkse et al. (1993) compared <strong>the</strong>ir collection<br />

with <strong>the</strong> holotype and noted <strong>the</strong> distinctive denticulate<br />

leaf border, composed <strong>of</strong> long narrow cells, running<br />

nearly to <strong>the</strong> leaf apex.<br />

156. Tortella limosella is known only from <strong>the</strong> Scottish<br />

type-locality, and has not been refound since its<br />

discovery in 1906.<br />

157. Tortella cirrifolia was synonymized with T. nitida by<br />

Sjögren (2001).<br />

158. Molecular studies by Werner et al. (2005b) show that<br />

<strong>the</strong> genus Trichostomum as listed here is heterogeneous,<br />

with T. brachydontium and T. crispulum nested<br />

within Weissia and T. triumphans very close to<br />

Pottiopsis caespitosa. Trichostomum connivens<br />

(Lindb. ex Broth.) Paris is recorded from <strong>the</strong><br />

Caucasus but is not known from Europe.<br />

Trichostomum contortum (Kunze) Sérgio is probably<br />

a synonym <strong>of</strong> T. brachydontium, although Sérgio<br />

(1985) thought that it was a good species.<br />

159. For synonymy <strong>of</strong> Hyophila treleasei, see Sollman<br />

(1993).<br />

160. Weissia leptocarpa Schimp. ex Besch. is treated by<br />

Sérgio & Carvalho (2003) as a doubtful species, but is<br />

accepted for Turkey by Kürschner & Erdag˘ (2005), in<br />

spite <strong>of</strong> <strong>the</strong> fact that Frey & Kürschner (1991) had<br />

treated it as doubtful and illegitimate.<br />

161. Weissia mittenii (Bruch & Schimp.) Mitt. is omitted,<br />

because it is designated above as a hybrid<br />

W.6mittenii (Bruch & Schimp.) Mitt. emend.<br />

A.J.E.Sm. Smith (pers. comm. to Hill) notes that W.<br />

mittenii has malformed capsules and abnormal spores.<br />

Following Werner et al. (2005b), Astomum is included<br />

in Weissia.<br />

162. A molecular study by Werner et al. (2004b) supports<br />

<strong>the</strong> species status <strong>of</strong> Weissia wimmeriana.<br />

163. Acaulon mediterraneum is treated by many authors as<br />

a variety <strong>of</strong> A. muticum, to which it is undoubtedly<br />

close.<br />

164. Aloina humilis was described from <strong>the</strong> Canary Islands<br />

by Gallego et al. (1998).<br />

165. Aloina obliquifolia is accorded species rank following<br />

Gallego et al. (1999).<br />

166. The Macaronesian Barbula semilimbata Dixon &<br />

Luisier was listed by Eggers (1982) but is a poorly<br />

known taxon, not recorded since it was described.<br />

167. Barbula convoluta var. sardoa is treated by some at<br />

species rank. Frahm & Ahmed (2004a) suggested <strong>the</strong><br />

name B. sardoa. However, <strong>the</strong> epi<strong>the</strong>t sardoa is correct<br />

at <strong>the</strong> varietal level but superfluous at <strong>the</strong> species level.

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