Bryological Monograph An annotated checklist of the mosses of ...

autoicous specimens and the differences he reported
from B. lonchocaulon (other than in sexuality) appear
inconstant in material from Britain.
295. Bryum bimum may merit species rank since it differs
from B. pseudotriquetrum in being synoicous rather
than autoicous, perhaps in having twice as many
chromosomes (Smith, 2004), and in average dimensions
of the leaf lamina cells and thickness of their cell
walls (Demaret & Empain, 1985; Zolotov, 2000).
However, they apparently do not differ much, if at all,
in ranges, frequency or ecology, and preliminary
molecular data (Hedenäs, pers. comm. to Holyoak)
suggest they are very close.
296. Bryum salinum differs from B. archangelicum in only a
few characters; a few intermediate specimens are
known.
297. Bryum schleicheri author citation follows Geissler
(1985).
298. Wijk et al. (1959) cited the original description of
Bryum torquescens as Bruch ex De Not., Syllab.
Musc., no. 163, 1838. Syed (1973) found this reference
to be wrong and he discounted ‘Fl. Sard. Exs., 1828’,
given by other authors because he suspected it is a
nomen nudum. The original description is therefore
taken as that by Bruch & Schimper (Bryol. Eur., 4 p.
119, pl. 20, 1839 (fasc. 6–9, Mon., p. 49, pl. 20)) partly
because this was the earliest description found by Syed
and partly because they refer to it as ‘Diese neue Art’.
299. Arts, Crundwell & Whitehouse (1995) showed that
Bryum pyriferum described from Tenerife is conspecific
with B. valparaisense, which was originally
described from Chile but is now also known from N.
America, Africa and Portugal.
300. Hedwig (1801) used the spelling zierii twice, but the
name Bryum zierii commemorates John Zier as was
pointed out by Crum & Anderson (1981). More recent
authors who amend it to zieri are therefore correcting
it according to Article 60.11 of the International Code
(Greuter et al., 2000).
301. Touw (1984) gave reasons for rejecting the name
Rhodobryum spathulatum in favour of R. ontariense.
302. The large family Mniaceae of Goffinet & Buck (2004)
is here divided following Koponen (1988) into the
three families traditionally included in Mniaceae and
the remainder, for which the only available name is
Mielichhoferiaceae.
303. Mielichhoferia elongata is often treated as a synonym
or variety of M. mielichhoferiana, but according to
Shaw (2000a), it is in reality a cryptic species.
304. Pohlia ramannii Warnst. is recorded from the territory
that was formerly in Finland and now in Russia
(Söderström, 1998). It is not mentioned by Ignatov &
Afonina (1992) and is treated here as doubtful.
Webera luisieri Dixon and W. maderensis Dixon &
Luisier, listed by Eggers (1982), are obscure taxa
whose identity is uncertain.
MOSSES OF EUROPE AND MACARONESIA 239
305. Sections of Pohlia are based on Shaw (1984), except
that we follow Smith (2004) in placing P. crudoides in
section Pohlia and P. flexuosa in section Cacodon.
306. Pohlia bolanderi was found in the Sierra Nevada, new
for Europe (Rams et al., 2004).
307. Pohlia ambigua was described from Austria but
remains a poorly known taxon. According to Shaw
(1982) it is a form of P. elongata. It is autoicous and is
therefore assigned to var. acuminata.
308. Species limits in the Pohlia nutans group require
further elucidation. Köckinger, Kučera & Stebel
(2005) studied P. schimperi and established that the
only really stable character separating it from P.
nutans is its red coloration; they accordingly gave it
subspecies rank and synonymized it with P. nutans
var. purpurascens. Pohlia marchica Osterwald was
described from near Berlin and subsequently collected
on Bornholm. It has not been found for many years,
and is signified by Nyholm (1993) as very close to P.
schimperi. Pohlia turonensis J.M.Couderc & Guédès,
nom. inval. was described from France. It is a close
relative of P. nutans and has not subsequently been
validated.
309. Pohlia saprophila, a mainly Asian species, is reported
from the Komi Republic, Russia (Zheleznova, 1994).
It may prove to be a dioicous race of P. elongata, but
needs further study.
310. For the synonymy of Pohlia flexuosa see Townsend
(1995).
311. Smith (2004) recognized two varieties of Pohlia
flexuosa in Europe, namely var. flexuosa and var.
pseudomuyldermansii nom. nud. (P. muyldermansii
var. pseudomuyldermansii Arts, Nordhorn-Richter &
A.J.E.Sm.). Townsend (1995) had noted that there is
continuous variation in bulbil morphology in Nepal
and India, and argued that the varieties should
therefore be treated as synonyms. He hypothesized
that there might be only two clones in Europe, with
var. muyldermansii possibly a recent introduction.
However Smith (pers. comm. to Hill) is of the opinion
that in Europe there has been differentiation into two
morphologically and ecologically distinct groups of
plants. In view of the distribution of var. pseudomuyldermansii
(Austria, British Isles and Switzerland)
it seems unlikely that the populations concerned
represent a single clone. The habitat of the varieties
appears to be markedly different in Europe. For these
reasons, Smith (above) makes the new combination P.
flexuosa var. pseudomuyldermansii.
312. Pohlia tundrae is reported from Central Europe (Düll,
1991, 1992; Müller, 2004).
313. Mnium lycopodioides is treated here as a full synonym
of M. ambiguum following Koponen (1994).
314. There are a number of records of Trachycystis
ussuriensis in the Russian Caucasus, e.g. Kharzivov
et al. (2004).