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The benthic ecology and community structure in Lyttelton Harbour ...

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51.<br />

SECTION 4<br />

COMMUNITIES<br />

4.1 Introduction<br />

Assemblages of animals are very· rarely present<br />

as discrete groups with clear cut boundaries. Group<strong>in</strong>g<br />

therefore must be <strong>in</strong>ferred from consideration of the<br />

<strong>in</strong>teraction of the fauna with variotis physical <strong>and</strong><br />

chemical factors. <strong>The</strong> habitat must be shown to be<br />

reasonably constant, support<strong>in</strong>g a stable population<br />

before the existence of a <strong>community</strong> is considered. In<br />

most regions one or more species seem to dom<strong>in</strong>ate samples,<br />

<strong>and</strong> provid<strong>in</strong>g they are a permanent feature of the local<br />

fauna, may be termed <strong>in</strong>dicator species. Such species<br />

usually have vary<strong>in</strong>g tolerances with<strong>in</strong> the study' zone<br />

<strong>and</strong> may serve to delimit communities. This approach may<br />

not be valid where a rarer animal shows a very patchy<br />

distribution at po<strong>in</strong>ts where its' narrow requirements<br />

are met· exactly.<br />

4.2 Community concepts<br />

<strong>The</strong> pioneer <strong>in</strong> <strong>benthic</strong> <strong>community</strong> <strong>ecology</strong> was<br />

undoubtedly Petersen whose major publications '(Petersen,<br />

1913 <strong>and</strong> 1914), set up basic pr<strong>in</strong>ciples which have been<br />

used by most later <strong>in</strong>ves gators as a start<strong>in</strong>g po<strong>in</strong>t<br />

for their own approaches. S<strong>in</strong>ce that time the recognition<br />

of communities have become a basic aim of <strong>benthic</strong><br />

ecological studies. Two schools of thought have evolved,<br />

the statistical <strong>and</strong> biological approaches. Petersen<br />

was an adherent of the former with the aim of avoid<strong>in</strong>g<br />

subjective conclusions. Conclusions reached by his

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