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The Cucurbitaceae of India: Accepted names, synonyms ...

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<strong>The</strong> <strong>Cucurbitaceae</strong> <strong>of</strong> <strong>India</strong> 59<br />

Checklist<br />

1. Actinostemma tenerum Griff., J. Asiat. Soc. Bengal 23(7): 643–644. 1854.<br />

Syntypes: <strong>India</strong>, Meghalaya, Khasia Hills, Griffith 2523 (K, W); <strong>India</strong>, Sadiya, upper<br />

Assam, also on Khasia Hills, T.E. Cantor s.n. (K).<br />

Distribution in <strong>India</strong>: Arunachal Pradesh, Assam, Bihar, Meghalaya, Mizoram, Uttar<br />

Pradesh, West Bengal.<br />

Distribution outside <strong>India</strong>: Bangladesh, Vietnam, Laos, Cambodia, Russia, China,<br />

Taiwan, Korea, and Japan (Schaefer and Renner, 2011a).<br />

Images: Griffith syntype: http://herbarium.univie.ac.at/database/detail.php?ID=63181<br />

http://apps.kew.org/herbcat/getImage.do?imageBarcode=K000742924<br />

GenBank: Sequences from Kocyan et al. (2007), e.g., DQ491007, DQ469135.<br />

Comments: Actinostemma comprises two other species, both in China. Based on genetic<br />

data, this is an isolated ancient lineage (Schaefer and Renner, 2011a, b; see also<br />

our Fig. 1). Ali Khan (2002) discusses the species’ occurrence in Uttar Pradesh.<br />

2. Benincasa fistulosa (Stocks) H.Schaef. & S.S.Renner, Taxon 60: 133. 2011.<br />

Citrullus fistulosus Stocks, Hooker’s J. Bot. Kew Gard. Misc. 3: 74, t. 3. 1851.<br />

Citrullus vulgaris Schrad. ex Eckl. & Zeyh. var. fistulosus (Stocks) J.L.Stewart, Punjab<br />

Pl. 96. 1869.<br />

Praecitrullus fistulosus (Stocks) Pangalo, Bot. Zhurn. S.S.S.R. 29: 203. 1944.<br />

Colocynthis citrullus (L.) Kuntze var. fistulosus (Stocks) Chakrav., Rec. Bot. Surv. <strong>India</strong><br />

17(1): 116. 1959.<br />

Citrullus lanatus (Thunb.) Matsum. & Nakai var. fistulosus (J.L.Steward) Babu, Herb.<br />

Fl. Dehra Dun 194. 1977, invalid name because Babu erred in the basionym he<br />

cited.<br />

Citrullus lanatus (Thunb.) Matsum. & Nakai var. fistulosus (Stocks) Chakrav., Fasc. Fl.<br />

<strong>India</strong> 11: 23. 1982, nom. illeg. isonym.<br />

Type: Pakistan [<strong>India</strong>], Kurrachee, Sinde, 1 Sep. 1850, Stocks s.n. (K).<br />

Distribution in <strong>India</strong>: Punjab, Rajasthan, Uttar Pradesh.<br />

Distribution outside <strong>India</strong>: Introduced (?) in tropical Africa.<br />

GenBank: Sequences from Dane and Lang (2004) and Kocyan et al. (2007), e.g.,<br />

DQ536719, DQ648185, AY522525.<br />

Comments: Cultivated in <strong>India</strong> and Pakistan as a vegetable. <strong>The</strong> origin <strong>of</strong> Benincasa<br />

fistulosa is unclear, and the species is currently only known in cultivation.

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