14ARACEAE10. Rhaphidophora dulongensis H. Li, Acta Bot. Yunnan.,Suppl. 5: 7. 1992.独 龙 崖 角 藤 du long ya jiao tengLianas. Stem green, cylindric, 1–2 m, 2–3 cm in diam.,internodes 0.5–2.5 cm. Petiole 41–49 × ca. 1.5 cm, basal 3–4cm sheathing; pulvinus indistinct; leaf blade pale green abaxially,green adaxially, oblong-ovate in outline, ca. 50 × 35 cm,subleathery, base rounded or subcordate, pinnately divided; pinnae6–9 per side, broadly linear, oblique, those at middle of leaf4–5 cm wide, apex truncate with falcate tip; costae 3 or 4,equally strong, parallel; sinuses nearly reaching midrib. Pedunclesolitary, cylindric, 15–20 cm × 8–10 mm. Spathe unknown.Spadix sessile, pale green, cylindric, ca. 10 × 1.8 cm. Ovaryquadrangular-cylindric or pentagonal-cylindric, apex truncate;stigma capitate. Fr. Nov–Mar.● Valley forests; below 2500 m. Yunnan (Gongshan).11. Rhaphidophora crassicaulis Engler & K. Krause inEngler, Pflanzenr. 37(IV. 23B): 52. 1908.粗 茎 崖 角 藤 cu jing ya jiao tengLianas. Branches dorsiventrally compressed, to 3 cm indiam., internodes short. Leaf blade ovate in outline, 30–55 ×22–36 cm, leathery, base obtuse, apex acute, pinnately divided;pinnae 6–10 or more per side, reaching midrib or nearly so,linear-falcate, base decurrent, apex truncate; primary lateralveins usually 2–4, equally strong, parallel, along midrib withperforations (8–20 × 2–5 mm), pinnae at middle of leaf 15–21 ×1–5 cm. Inflorescences terminal on lateral branches, solitary,erect; peduncle robust, cylindric, 12–20 × 1–1.6 cm. Spatheunknown. Spadix sessile, white, cylindric, 10–17 cm, 2–2.5 cmin diam. at base. Ovary cylindric, 6–8 × ca. 2 mm; ovules many,oblong, funicle long; style broadly conic; stigma circular. Fr.Nov–Dec.Dense tropical forests, on trees or over stones; below 1300 m.Guangxi, Hainan, Yunnan [N Laos, N Vietnam].12. Rhaphidophora decursiva (Roxburgh) Schott, Bonplandia(Hannover) 5: 45. 1857.爬 树 龙 pa shu longPothos decursivus Roxburgh, Fl. Ind. 1: 456. 1820 [“decursiva”];Monstera decursiva (Roxburgh) Schott; Rhaphidophoraaffinis Schott; R. eximia Schott; R. grandis Schott; R.insignis Schott; Scindapsus decursivus (Roxburgh) Schott.Lianas, very large, to 20 m or more, very robust. Stemgreen at back, yellow at front, robust, 3–5 cm in diam., rooting,internodes 1–2 cm. Leaves scattered, each foliage leaf separatedfrom next by several (up to 7) cataphyll-bearing nodes; petiolegreen, cylindric, 20–40 cm, shallowly canaliculate; pulvinusindistinct; petiolar sheath extending to ca. 2/3 along petiole;juvenile leaf blade orbicular, ca. 16 × 13 cm, margin entire,apex abruptly acute; mature leaf blade pale green abaxially,green adaxially, oblong-ovate, ovate in outline, 60–70[–100] ×40–50 cm, base subcordate, margin irregularly and asymmetricallypinnately divided, apex acute; pinnae 9–15 per side, ca. 20× 3–5 cm at mid-leaf, base narrowed, apex truncate with ascendingfalcate tip, with a strong costa. Inflorescences axillary,solitary; peduncle green, cylindric, very robust, 11–20 × 1.5–3cm. Spathe initially involute, afterward spreading, yellow onboth sides, cymbiform, oblong-ovate, 17–20 × 10–12 cm.Spadix sessile, gray-green, cylindric, 15–16 × 2–3 cm, baseoblique. Filaments flat, ca. 5 mm, nearly as long as ovary, baseangustate; anthers yellow, oblong, less than 1 mm. Ovary hexagonal-conic,ca. 5 × 3 mm; style distinct, ca. 1 cm; stigmayellow, longitudinally oblong; ovules many; funicle laterallyinserted, slender. Infructescence 15–20 × 5–5.5 cm. Berrygreen-white, base white or yellow, hexagonal-obconic, ca. 18 ×5 mm, apex with a persistent style, stylar region green-white,sloughing to reveal white or yellow pulp cavity. Fl. May–Aug,fr. mature next Jul–Sep.Monsoon rain forests, valley evergreen broad-leaved forests,creeping on ground, over rocks, or climbing against trees; below 2200m. Fujian, Guangdong, Guangxi, Guizhou, Hainan, Sichuan, Taiwan,SE Xizang (Mêdog, Zayü), Yunnan [Bangladesh, Bhutan, Cambodia,NE India, Laos, Myanmar, Nepal, Sri Lanka, N Thailand, Vietnam].The stems and leaves are used medicinally for treating traumaticinjuries, fractures, swellings, colds, lumbago, snake bites, coughs, andbronchitis.There has been much confusion over Rhaphidophora decursivaand Epipremnum pinnatum because both species have large, pinnatifidleaves, but E. pinnatum has a leaf blade with tiny perforations (ca. 2mm) along and adjacent to the midrib, an ovary with 2 or 3 ovules, anda sessile, longitudinally linear-oblong stigma. There is also a differencein leaf arrangement: R. decursiva has foliage leaves scattered and interspersedwith several (up to 7) cataphyll-bearing nodes, whereas E. pinnatumhas leaves often clustered distally without such cataphyll-bearingnodes.7. EPIPREMNUM Schott, Bonplandia (Hannover) 5: 45. 1857.麒 麟 叶 属 qi lin ye shuLi Heng ( 李 恒 Li Hen); Peter C. BoyceLianas, large to very large. Stems robust, woody, branched and rooting. Leaves long petiolate; petiole with pulvinus at junctionwith leaf blade, somewhat sheathing at base; leaf blade large, entire, pinnatifid, or pinnatisect, with perforations along midrib or not.Inflorescences terminal or sometimes displaced, solitary; peduncle robust. Spathe caducous, cymbiform, ovate, acute or acuminate.Spadix sessile. Flowers bisexual, sometimes with female ones at spadix base and these sterile, lacking perigone. Stamens 4(–6); filamentslinear, flat; anthers much shorter than filaments; thecae ellipsoidal, longitudinally dehiscent. Ovary polygonal-cylindric, apextruncate, 1-loculed; ovules 2–4, inserted at base of parietal placenta, sometimes 6–8, distichous on parietal placentae, anatropous,with short funicle; stigma sessile, linear-oblong, longitudinal. Fruit small. Seeds reniform, usually striate.About 20 species: tropical Asia, Australia, Pacific islands; one species in China.Epipremnum aureum (Linden & Andre) Bunting is cultivated in gardens.
ARACEAE 151. Epipremnum pinnatum (Linnaeus) Engler, Pflanzenr.37(IV. 23B): 60. 1908.麒 麟 叶 qi lin yePothos pinnatus Linnaeus, Sp. Pl., ed. 2, 2: 1324. 1763;Epipremnum angustilobum K. Krause; E. elegans Engler; E.formosanum Hayata; E. glaucicephalum Elmer; E. merrillii(Engler) Engler & K. Krause; E. mirabile Schott; Monsteracaudata (Roxburgh) Schott; M. dilacerata (K. Koch & Sello)C. Koch; M. pinnata (Linnaeus) Schott; M. pinnatifida (Roxburgh)Schott; Polypodium laciniatum N. L. Burman; Pothoscaudatus Roxburgh; P. pinnatifidus Roxburgh; Rhaphidophoracaudata (Roxburgh) Schott; R. cunninghamii Schott; R. dilacerata(K. Koch & Sello) C. Koch; R. formosana Engler (1879);R. formosana (Hayata) M. Hotta (1970); R. laciniata (N. L.Burman) Merrill; R. lovellae F. M. Bailey; R. merrillii Engler;R. neocaledonica Guillaumin; R. pertusa (Roxburgh) Schottvar. vitiensis (Schott) Engler; R. pinnata (Linnaeus) Schott; R.pinnatifida (Roxburgh) Schott; R. vitiensis Schott; R. wallichiiSchott; Scindapsus caudatus (Roxburgh) Schott & Endlicher; S.dilaceratus K. Koch & Sello; S. forsteri Endlicher; S. pinnatifidus(Roxburgh) Schott; S. pinnatus (Linnaeus) Schott;Tornelia dilacerata (K. Koch & Sello) Schott.Root climbers, to 15 m. Mature stem 5–40 mm in diam.,internodes separated by prominent leaf scars, obscured bysheath fibers, active stems lustrous green with prominent irregularlongitudinal whitish crests. Foliage leaves often clustereddistally; petiole dark green, 19.5–60 cm × 3–13 mm, canaliculate,smooth; apical pulvinus 16–70 × 3–5 mm, basal pulvinus3–7 × 1–1.5 cm; petiolar sheath extending to halfway alongapical pulvinus, disintegrating into netted fibers; leaf bladeovate to oblong-elliptic, 10–93 × 5–60 cm, submembranous,base slightly cordate, regularly pinnatifid to pinnatisect, apexacuminate; pinnae paler abaxially, adaxially dark green, 1.2–6.5cm wide, with tiny perforations along and adjacent to midrib,pinnae each with 1 primary lateral vein and several parallelinterprimary veins, higher order venation reticulate, midribprominently raised abaxially, impressed adaxially. Inflorescenceseveral together, partially obscured by netted fibers; pedunclepale green, 5.5–21.5 cm × 4–10 mm, terete. Spathe dull yellowto mid-green outside, dull yellow or pale green inside, canoeshaped,7–23.5 × 3–15 cm, stoutly attenuate to 15 mm, openingalmost flat at anthesis. Spadix sessile, white, later glaucousgray-green to yellow, cylindric, 8.5–25 × 1.1–3.5 cm, baseslightly obliquely inserted. Flowers 3–7 mm in diam. Ovarycylindric, 4–12 × 2–7 mm, basal part slightly compressed;ovules 2 or 3; stylar region trapezoid, 3–7 × 1.5–4 mm, ratherrobust, truncate; stigma longitudinally linear-oblong, 2–6 × 0.1–0.5 mm. Fruit mid-green. Seeds reniform, ca. 4.5 × 3.5 mm. Fl.Apr–Jul.Tropical rain forests, mountain forests, creeping and climbing ontrees, rocks, or stone walls; below 2000 m. Guangdong, Guangxi, Hainan,Taiwan, Yunnan [Bangladesh, Cambodia, India, Indonesia, Japan,Laos, Malaysia, Myanmar, Papua New Guinea, Philippines, Singapore,Thailand, Vietnam; Australia (Queensland), Pacific islands].The plants are used medicinally to treat traumatic injuries, abscesses,and rheumatic arthralgia.8. SCINDAPSUS Schott in Schott & Endlicher, Melet. Bot. 21. 1832.藤 芋 属 teng yu shuLi Heng ( 李 恒 Li Hen); Peter C. BoyceLianas, creeping and climbing against trees with aid of adhesive roots. Stem robust, branched. Leaves spiro-distichous; petioleusually sheathing or winged throughout its length; leaf blade undivided, pinnately veined, with very numerous parallel, thin lateralveins. Inflorescences solitary, near stem apices; peduncle at last laterally protruding from petiolar sheath. Spathe not constricted,initially involute and tubular, afterward somewhat inflated, finally expanding only by a rather narrow longitudinal slit, never entirelyor conspicuously flat-spreading, falling off as a whole leaving a thick scar on apex of peduncle. Spadix drying black, cylindric, muchthicker than peduncle. Flowers many, bisexual, naked. Stamens 4; filaments strap-shaped. Ovary with flat, often subrhomboid orquadrangular apex, 4–6-angular, 1-loculed; ovule 1, basal; stigma sessile, disciform to linear. Seeds subreniform.About 36 species: tropical Asia; one species in China.1. Scindapsus maclurei (Merrill) Merrill & F. P. Metcalf,Lingnan Sci. J. 21: 5. 1945.海 南 藤 芋 hai nan teng yuRhaphidophora maclurei Merrill, Philipp. J. Sci. 21: 337.1922; Scindapsus megaphyllus Merrill.Lianas. Stem robust, ca. 2.5 cm in diam. Petiole 26–32 cm,base imbricate, sheath reaching pulvinus; leaf blade pale greenabaxially, green adaxially, drying pale brown, oblong-elliptic orovate-elliptic, 23–46 × 12–24 cm, leathery, base subcordate,margin entire, apex acute or shortly acuminate; lateral veinsnumerous, diverging at 70–80° from midrib. Peduncle robust.Spathe yellow, involute-tubular, 18–22 cm, apex acuminate.Spadix sessile, cylindric, ca. 15 × 2.5 cm. Filaments ca. 4 mm;anthers ovoid, ca. 3 mm. Fruit ca. 9 × 5 mm, apex truncate, 1-loculed, 1-seeded. Fl. Nov–Dec.Dense tropical forests, creeping and climbing on trees and stonewalls; 400–600 m. Hainan [N Thailand, N Vietnam].Scindapsus maclurei differs from the vegetatively very similarRhaphidophora megaphylla in leaf venation (primary lateral veins numerousand not differentiated from the interprimary veins, diverging at70–80° from midrib in S. maclurei vs. primary lateral veins 10–13 oneach side, diverging at 80–90° from midrib, and strongly differentiatedfrom the interprimary veins in R. megaphylla).
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ARACEAE 651a. Spathe throat margin
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ARACEAE 67stripes above neuter flow
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ARACEAE 69narrower, with abundant a
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ARACEAE 71Spadix up to 2.3 cm; fema
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ARACEAE 7324. COLOCASIA Schott in S
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ARACEAE 75sessile or subsessile, di
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ARACEAE 77reflexed, then deliquesce
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ARACEAE 79florescences solitary or