78ARACEAEPachycaul herbs, massive, to 4 m, evergreen, latex slightlymilky. Stem erect, to ca. 1.5 m, decumbent. Leaves severaltogether, clustered at tips of stems in larger plants; petiole to 1.3m, sheathing in proximal 1/3–1/2; leaf blade light green on bothsurfaces, ovate-sagittate, bluntly triangular, up to 120 × 50 cm,± erect, margin entire to very slightly sinuous; posterior lobes1/3–1/2 length of anterior, somewhat rotund, often overlapping,naked in sinus in mature plants, weakly peltate in juveniles;primary lateral veins ca. 9 on each side of anterior lobe, axillaryglands distinct, secondary venation not forming interprimarycollective veins. Inflorescences paired among leaf bases, subtendedby membranous cataphylls; peduncle barely exceedingcataphylls at anthesis. Spathe 13–35 cm, constricted ca. 1/6from base; proximal spathe green, ovoid; limb cowl-like atanthesis, later reflexed, then deliquescent, pale yellow, broadlyoblong-lanceolate, 10.5–29 cm. Spadix slightly shorter thanspathe, shortly stipitate; female zone conic-cylindric, 1–2 × ca.1.5 cm; pistil pale green, ca. 3 mm in diam.; stigma yellow,sessile, 3–5-lobed; sterile zone slightly equaling female zone,whitish; synandrodes rhombic-hexagonal, ca. 2.5 mm in diam.;male zone whitish, cylindric, 3–7 × ca. 2 cm; synandria 5–9-merous, rhombic-hexagonal, convex-topped, ca. 2 mm in diam.;appendix yellowish, slightly tapering, at least 1/2 length ofspadix. Fruiting spathe green, oblong-ellipsoid, ca. 8 cm. Fruitripening scarlet, ellipsoid, ca. 12 × 8 mm.Ditches and wet areas of farmlands and wastelands, never awayfrom human disturbance; sea-level to 800 m. S Fujian, Guangdong,Guangxi, Guizhou, Hainan, S Sichuan, Taiwan, S Xizang, Yunnan [originallyfrom tropical Asia; now pantropical].It is not clear where, if anywhere, Alocasia macrorrhizos occurswild. It has evidently been prehistorically distributed widely in tropicalAsia as a subsistence crop and is now pantropical by introduction as anornamental.5. Alocasia cucullata (Loureiro) G. Don in Sweet, Hort. Brit.,ed. 3, 631. 1839.尖 尾 芋 jian wei yuArum cucullatum Loureiro, Fl. Cochinch. 2: 536. 1790;Alocasia rugosa (Desfontaines) Schott; Caladium cucullatum(Loureiro) Persoon; C. rugosum Desfontaines; Colocasiacochleata Miquel; C. cucullata (Loureiro) Schott; C. rugosa(Desfontaines) Kunth; Panzhuyuia omeiensis Z. Y. Zhu.Herbs, clumping, small to medium sized, somewhat robust,to 1 m, evergreen. Stems erect, hypogeal, basally muchbranched. Leaves many together; petiole weakly D-shaped incross section, 25–30(–80) cm, sheath reaching to ca. 1/2 way,margins membranous; leaf blade broadly ovate-cordate, 10–40× 7–28 cm, base shallowly cordate, apex acute; primary veins 4on each side, radiating from petiole, arching, interprimary veinsnot forming a collective vein. Inflorescences rarely produced,usually solitary, sometimes paired, among leaf bases, subtendedby membranous cataphylls; peduncle 20–30 cm. Spathe green,9–15 cm; proximal spathe 4–8 × ca. 2.5 cm; limb narrowlycymbiform, 5–10 × 3–5 cm. Spadix 8–14 cm; female zonecylindric, 1.5–2.5 cm × ca. 7 mm; sterile zone 2–3 cm × ca. 3mm; male zone yellow, ca. 3.4 cm × 8 mm; appendix yellowish,narrowly conic, ca. 3.5 cm × 5 mm. Fruit rarely produced, asubglobose berry, 6–8 mm in diam., ripening red. Fl. May.Watersides, by fields, ?wild or cultivated; below 2000 m. Fujian,Guangdong, Guangxi, Guizhou, Hainan, Sichuan, Taiwan, Yunnan[Bangladesh, NE India (Khasi Hills, Sikkim), Laos, Myanmar, Nepal,Sri Lanka, Thailand, Vietnam].The plants are used externally for detoxifying viper bites and arealso used for treating abscesses, rheumatism, and arthritis. It is an importantgood-luck plant in Buddhist temples in Laos and Thailand.6. Alocasia hypnosa J. T. Yin, Y. H. Wang & Z. F. Xu, Ann.Bot. Fenn. 42: 395. 2005.紫 苞 海 芋 zi bao hai yuHerbs, terrestrial or lithophytic, to 100 cm tall, seasonallydormant. Stem erect or a tuberlike rhizome; rhizome ca. 10 ×13.5 cm; stolons numerous per plant, pale green, simple,trailing horizontally or pendent, 56–110 cm, internodes cylindric,ca. 5.5 × up to 1 cm, with light green cataphylls, tuberclesproduced terminally, to 4 × 3 cm. Leaves 3–6 together;petiole light greenish, glossy, terete, to 104 × 3–7.5 cm, sheathto 50 cm, membranous; leaf blade triangular-sagittate, to 82 ×64 cm; basal lobes naked in sinus, membranous, glossy brightgreen; primary lateral veins 8 per side, conspicuous, interprimaryveins forming a feeble collective vein. Inflorescences 2 or3 together, appearing with leaves; peduncle pale green, cylindric,to 90 cm. Spathe to 28 cm, not constricted between basalconvolute part and limb; proximal spathe green, fusiform, ca.6.5 × 3 cm; limb arched, erect at anthesis, later flopping forward,then deliquescent, purple-pink or whitish, oblong-lanceolate,to 24 × 20 cm. Spadix sessile, shorter than spathe; femalezone cylindric, ca. 1.5 × 2 cm; pistil oblong, ca. 5 mm; styleshort; stigma 3- or 4-lobed; sterile zone cylindric, ca. 5.5 × 1–1.5 cm; synandrodes depressed oblong to depressed ovate, apextruncate or concave; male zone white, cylindric, ca. 3.5 × 2 cm;synandria 5- or 6-merous, rhombic-hexagonal, truncate; appendixwhite, narrowly conic, to 16.5 × 2.5 cm. Fruiting spatheellipsoid, ca. 3 × 2.5 cm. Fruit ellipsoid, ca. 1.5 × 0.5 cm,ripening scarlet.Humid forest margins and exposed areas of forest regrowth inlimestone mountains; 900–1000 m. SW Yunnan (Menglian) [N Laos, NThailand].7. Alocasia longiloba Miquel, Fl. Ned. Ind. 3: 207. 1856.尖 叶 海 芋 jian ye hai yuAlocasia amabilis W. Bull; A. cochinchinensis Pierre exEngler & K. Krause; A. cuspidata Engler; A. lowii Hooker var.veitchii (Lindley) Engler; A. veitchii (Lindley) Schott; Caladiumveitchii Lindley.Herbs, terrestrial, small to robust, to 150 cm tall, evergreenor seasonally dormant. Stem rhizomatous, elongate, erect to decumbent,sometimes subtuberous, 8–60 × 2–8 cm. Leaves 1–3together, subtended by lanceolate papery-membranous cataphylls;petiole terete, 30–120 cm, glabrous, sheathing in proximal1/4, purple-brown to green, strikingly obliquely mottledchocolate-brown; leaf blade pendent, green or flushed withpurple abaxially, dark green often with major venation graygreenadaxially, narrowly hastate-sagittate, 27–85 × 14–40 cm;posterior lobes peltate for (5–)10–30% of their length, acute,anterior lobe with 4–8 primary lateral veins on each side, axillaryglands conspicuous, interprimary collective veins weak. In-
ARACEAE 79florescences solitary or paired; peduncle 8–18 cm, resemblingpetioles, subtended by cataphylls. Spathe 7–17 cm, abruptlyconstricted 1.5–3.5 cm from base; proximal spathe green,ovoid; limb erect, later reflexing, pale green, lanceolate, canoeshaped,5.5–7.5 cm. Spadix somewhat shorter than spathe, 6–13cm, stipitate, stipe whitish, conic, ca. 5 mm; female zone 1–1.5cm; pistil green, subglobose, 1.5–2 mm in diam.; stigma white,subsessile or on a slender style to ca. 0.5 mm, 3- or 4-lobed;sterile zone 7–10 mm, narrower than fertile zones; synandrodesmostly rhombic-hexagonal, flat-topped; male zone ivory, subcylindric,tapered at base, 1.2–2.5 cm × 4.5–8 mm; synandria4–6-merous, ± hexagonal, ca. 2 mm in diam.; appendix paleorange, 3.5–9 cm, subcylindric, tapering to a point. Fruitingspathe glossy green, ovoid, 4–7 cm. Fruit ripening orange-red,globose-ellipsoid, ca. 1.5 × 0.75 cm. Fr. Aug–Oct.Tropical forests, thickets; 100–1000 m. Guangdong, Hainan, SYunnan [Cambodia, Indonesia, Laos, Malaysia, S Myanmar, Singapore,Thailand, Vietnam].8. Alocasia acuminata Schott, Bonplandia (Hannover) 7: 28.1859.越 境 海 芋 yue jing hai yuHerbs, terrestrial, small to medium sized, slightly robust,to 75 cm tall, evergreen. Stem rhizomatous, generally elongate,erect, later decumbent, 8–75 × 2–6 cm. Leaves to 5 together,subtended by lanceolate papery-membranous cataphylls; petiolebright green, 15–80 cm, glabrous, sheathing in proximal 1/4;leaf blade spreading, bright green, narrowly hastate-sagittate toovate-hastate, 15–60 × 8–20 cm; posterior lobes 1/4–1/3 lengthof anterior, peltate for 25–30% of their length, acute; anteriorlobe with 3–6 primary lateral veins on each side, axillary glandshardly conspicuous, interprimary collective veins weakly defined.Inflorescences usually solitary; peduncle green, 9–20cm, subtended by cataphylls. Spathe 7–10 cm, moderately constricted1.5–2.5 cm from base; proximal spathe green, ovoid;limb pale green, lanceolate, canoe-shaped, 5.5–7.5 cm. Spadixsubequaling spathe, 6–9.5 cm, sessile; female zone 1–1.5 cm;pistil green, subglobose, 1.5–2 mm in diam.; stigma white, subsessile,very slightly lobed; sterile zone 7–10 mm, narrowerthan fertile zones; synandrodes narrowly rhombic-hexagonal,flat-topped; male zone ivory, subcylindric, 1.2–2.5 cm × 4.5–8mm; synandria 4–6-merous, ± hexagonal, ca. 2 mm in diam.;appendix white, 2.5–3.5 cm, demarcated from male zone bya strong constriction, narrowly conic. Fruiting spathe green,ovoid, 3–4 cm. Fruit ripening orange-red, globose-ellipsoid, ca.0.75 cm in diam.Everwet to seasonal perhumid evergreen forests, usually in deepshade, less often on track sides; 600–1800 m. S Yunnan [Bangladesh,NE India, N Laos, N Myanmar, Nepal, N Thailand, ?N Vietnam].Apiospermum Klotzsch; Kodda-Pail Adanson.26. PISTIA Linnaeus, Sp. Pl. 2: 963. 1753.大 漂 属 da piao shuLi Heng ( 李 恒 Li Hen); Peter C. BoyceHerbs, small, free-floating, evergreen, stoloniferous, acaulescent, with spreading feathery roots. Leaves several in a rosette,densely pubescent; petiole very short, almost absent; sheath ligulate, very short, very thin, scarious at base; leaf blade obovate-cuneateto obovate-oblong, somewhat spongy, apex rounded, truncate, or retuse; midrib absent, primary veins subparallel, all arising frombase, diverging slightly and running into margin near apex, strongly prominent abaxially, higher order venation reticulate. Inflorescencesolitary, very small, much shorter than leaves; peduncle very short, pubescent. Spathe somewhat constricted centrally, pubescentoutside, glabrous inside, proximal margins connate with each other and with ovary wall forming tube, free margins between tubeand blade folded between stigma and male organs forming a partition between a distal male partial chamber and a proximal femaleone; limb erect, ovate, expanded, acute-acuminate. Spadix shorter than spathe, mostly adnate to spathe, only apical male zone free;female zone with single gynoecium at base and a thin, green, pouch-shaped flap just below spathe partition; male zone subtended bya thin, marginally lobed, green, annular flap, consisting of a basally naked spadix axis supporting a single whorl of 2–8 flowers,naked axis sometimes extending slightly beyond. Flowers unisexual, naked. Gynoecium obliquely adnate to spadix axis; ovaryovoid, 1-loculed; ovules numerous, orthotropous; placenta broad, apparently parietal, probably morphologically basal; stylar regionattenuate, bending inward toward male flowers; stigma disciform-subcapitate, small. Male flower a synandrium consisting of 2 connatestamens; thecae dehiscing by single apical slit. Fruit a thin-walled, utricular, several-seeded, ellipsoid berry, irregularly breakingup and decaying to release seeds. Seed barrel-shaped, ± subtruncate and excavated at apex and base; testa thick, reticulate-alveolate,thicker and with operculum at micropylar end; embryo obovoid to conic; endosperm copious.One species: widely distributed in tropical and subtropical regions, including China.1. Pistia stratiotes Linnaeus, Sp. Pl. 2: 963. 1753.大 漂 da piaoApiospermum obcordatum Klotzsch, nom. illeg. superfl.;Pistia obcordata Schleiden, nom. illeg. superfl.Morphological characters are the same as those of thegenus. Plants aquatic, floating. Roots many, pendulous in water,feathery. Leaves in rosettes, 1.3–10 × 1.5–6 cm. Spathe white,5–12 mm. Fl. May–Nov.Water fields, lakes, ponds. Fujian, Guangdong, Guangxi, Taiwan,Yunnan; cultivated in Anhui, Hubei, Hunan, Jiangsu, Jiangxi, Shandong,Sichuan [tropical and subtropical regions worldwide].The plants are cultivated for animal feed and are used to treatswelling and urinary tract infections.
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ARACEAE天 南 星 科 tian nan xin
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ARACEAE 523a. Female zone of spadix
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ARACEAE 7rolled, apex rounded to ac
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ARACEAE 9several to many, borne in
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ARACEAE 11reiterating and overtoppi
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ARACEAE 13cent; leaf blade oblong,
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ARACEAE 151. Epipremnum pinnatum (L
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ARACEAE 171. Calla palustris Linnae
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ARACEAE 1912. SCHISMATOGLOTTIS Zoll
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ARACEAE 21fall, the more aquatic on
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ARACEAE 23apical pore or reniform t
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ARACEAE 2510b. Appendix terete or d
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