36ARACEAEbase, rest naked; staminodes erect or spreading, yellowish,clavate, 6–7 mm; male zone ca. 5 mm; male flowers yellow;appendix erect, purple, cylindric, ca. 10 cm × 2 mm. Fl. lateApr–Aug. 2n = 10*.● Yunnan (Jinping).7. Typhonium roxburghii Schott, Aroideae, 12. 1855.金 慈 姑 jin ci guUnderground part a short, tuberous rhizome, subglobose orglobose. Petiole 10–35 cm; leaf blade yellowish green abaxially,dark green adaxially, reniform-hastate, triangular in outline,shallowly or deeply 3-lobed; terminal lobe ovate, (5–)9–17× 4–14 cm, acuminate; lateral lobes spreading, obliquely ovateor narrowly ovate. Inflorescence appearing after leaves; pedunclecreamy red with dark purple dots, 2–9 cm. Spathe outsidewhitish or pale green with purple striae, inside purple, baseovoid, 2–3.5 × ca. 1.5 cm; limb outside purplish mixed withdirty green flush or striae, or purple all over, inside deep richpurple, ovate-lanceolate, abruptly tapering from below middle,usually twisted at tip, 13–15 × ca. 5 cm. Spadix subequalingspathe; female zone 4–5.5 mm; ovary white or yellowish,ovoid; stigma purple, 1-loculed, 1-ovuled; sterile zone 1.5–2.2cm, proximal 5–7 mm densely covered with staminodes, restnaked; staminodes downcurved, yellow or pale yellow, subcylindric,8–9 mm, acute; male zone 7–12 mm; male flowersconsisting of 2 or 3 stamens, yellow; anthers sessile, openingwith apical pore; appendix dark purple, narrowly conic, 12–15cm, 3–4 mm in diam. at base, base truncate with a pale red stipeca. 2 mm. Berries pale green at first, white when mature, ovoidor ellipsoid, 1-seeded. Fl. and fr. May–Aug.Taiwan, Yunnan [Bangladesh, India, Indonesia, Japan (BoninIslands), Malaysia, New Guinea, Philippines, Sri Lanka, Thailand;introduced in E Africa, W Australia, and South America].8. Typhonium trifoliatum Wang & Lo ex H. Li, Y. Shiao & S.L. Tseng, Acta Phytotax. Sin. 15(2): 105. 1977.三 叶 犁 头 尖 san ye li tou jianUnderground part a short, tuberous rhizome, ca. 1.5 × 1.2cm. Petiole 6–12 cm; leaf blade rarely entire and then linear,usually deeply 3-lobed or 3-partite; segments sessile, linear;terminal segment 8–10 cm × 3–8 mm, acuminate; lateral segmentshorizontally spreading, 1.7–4.5 cm × 3–4 mm. Inflorescenceappearing after leaves; peduncle 8–10 cm. Spathe convoluteat base, dark purple, basally ovoid, ca. 1.5 × 1.2 cm,apex constricted; limb ovate-lanceolate, ca. 17 × 5–6 cm, distalhalf narrowed, linear, recurved. Spadix shorter than spathe, 13–17 cm; female zone ca. 5 mm; ovary ovoid, 1-loculed, 1-ovuled; stigma disciform; sterile zone ca. 1.4 cm, proximal 1–2mm covered with staminodes, rest naked; staminodes downcurved,filiform; male zone 8–18 mm; appendix shortly stipitate,suberect, 10–13 cm, base swollen, 3–5 mm in diam., subtruncate.Berries ovoid, 1-seeded. Fl. and fr. Jul–Aug.● Hebei, Nei Mongol, Shaanxi, Shanxi.9. Typhonium trilobatum (Linnaeus) Schott, Wiener Z. Kunst1829: 732. 1829.马 蹄 犁 头 尖 ma ti li tou jianArum trilobatum Linnaeus, Sp. Pl. 2: 965. 1753.Underground part a short, tuberous rhizome, subglobose orsubcylindric, producing few annual offsets or splitting up.Petiole green or variously flushed with purple, 25–40 cm; leafblade cordate-ovate in outline, usually deeply 3-lobed, rarely5-lobed; central lobe ovate, 10–15 × 6–11 cm, acuminate,sometimes mucronate; lateral lobes 8–13 cm. Inflorescenceappearing after leaves; peduncle 5–10 cm, elongating in fruit.Spathe convolute at base, outside green, inside green, to 30 cm,ovoid or ellipsoid, 2.5–3.5 × 1–1.5 cm, constricted at apex; limbspreading, outside green, inside dark purple to reddish purple,ovate-lanceolate, ca. 15 × 5–8 cm, apex acuminate. Spadixshorter than spathe; female zone slightly conic, 7–10 mm;ovary yellowish green; stigma sessile, dark or mid-purple,disciform; sterile zone 2–3 cm, proximal half densely coveredwith staminodes, distal half naked; staminodes strongly curledbut mostly directed downward and covering most of femalezone, filiform, 7–12 mm; male zone 1.5–2 cm; stamens pink;appendix shortly stipitate, glossy purple or reddish, narrowlyconic, 5–12 cm × 4–7 mm, base truncate, apex acute or subacute.Fruiting zone with spathe remaining; berries at first greenwith purple spots, white when mature, ellipsoid, 1- or 2-seeded.Fl. May–Jul.Tropical secondary forests, thickets, grasslands, roadsides; below700 m. Guangdong, Guangxi, Hainan, S Yunnan [Bangladesh, Bhutan,Cambodia, India, Laos, Malaysia, Myanmar, Nepal, Sri Lanka, Thailand;introduced in W Africa, W Borneo, Neotropics, Philippines, andSingapore].19. SAUROMATUM Schott in Schott & Endlicher, Melet. Bot. 17. 1832.斑 龙 芋 属 ban long yu shuLi Heng ( 李 恒 Li Hen); Wilbert L. A. HetterscheidHerbs, seasonally dormant. Underground part a depressed-globose tuber. Leaves entire, sagittate-hastate, 3-lobed or -partite topedatisect. Inflorescence simultaneous with or preceding leaves; peduncle short. Spathe divided into a connate or strongly convolutebasal part with a ± strong apical constriction and a spreading lanceolate to ovate-triangular limb. Spadix as long as spathe, with 4zones: a proximal female zone, an intermediate sterile zone with staminodes all over or only at base, a male zone, and a sessile orstipitate appendix. Female flowers 1-loculed with 1 or 2(or 3) orthotropous basal ovules. Proximal staminodes clavate, distal ones(when present) irregularly cristate or elongate aristate. Male flowers with 1–3 stamens opening by slits or pores; connective sometimesprolonged. Fruiting zone with spathe caducous or remaining; berries ovoid, 1(or 2)-seeded. Seeds with endosperm.Eight species: Bhutan, Cambodia, China, India, Indonesia, Laos, Myanmar, Nepal, Thailand, Vietnam; seven species (two endemic) in China.See the comment under Typhonium above.
ARACEAE 371a. Sterile zone between male and female zones largely smooth and terete or grooved, except for proximal partcarrying staminodes.2a. Leaf and outside of spathe densely hirsute .............................................................................................................. 5. S. hirsutum2b. Leaf and outside of spathe glabrous.3a. Spathe base fully closed (margins connate), limb inside with background off-white, almost entirelyhidden by numerous strongly confluent, oval, deep maroon spots; leaves always pedate .............................. 7. S. venosum3b. Spathe base convolute (margins overlapping), limb inside uniformly whitish green, or with purplestripes, or uniformly purple; leaves varying (often in one plant) from entire to pedate ........................... 2. S. diversifolium1b. Sterile zone between male and female zones fully covered with staminodes, distally gradually becomingsmaller.4a. Leaves simple, cordate, at least 10 cm ................................................................................................................. 4. S. giganteum4b. Leaves pedate.5a. Spathe base convolute ..................................................................................................................................... 6. S. horsfieldii5b. Spathe base fully closed, margins connate.6a. Proximal staminodes gradually changing upward from clavate through irregularly short andcurved to irregularly cristate or strongly reduced to almost absent; petiole often with 1 or afew intercalary bulbils near base ....................................................................................................... 3. S. gaoligongense6b. Intermediate staminodes absent between proximal (clavate) and distal (irregularly cristate)staminodes ................................................................................................................................................... 1. S. brevipes1. Sauromatum brevipes (J. D. Hooker) N. E. Brown, Gard.Chron., ser. 3, 34(2): 93. 1903.短 柄 斑 龙 芋 duan bing ban long yuTyphonium brevipes J. D. Hooker, Fl. Brit. India 6: 511.1893.Tuber globose, 1–1.5 cm in diam., producing numeroustiny, annual bulbils. Petiole pinkish or dull purplish red, 15–30cm × ca. 2 mm; leaf blade green adaxially, 5–9-pedatifid; lobes± fused at base; central lobe linear-lanceolate or narrowlyelliptic, 6.5–25 × 1–5 cm, margin undulate, with or without apurple midrib; outermost lobes progressively smaller. Inflorescence(s)appearing alongside developing leaves; peduncle palegreen, 2–2.5 cm. Spathe outside base ocher or yellow-brownwith purple spots or faint pinkish spots, inside ± cream-coloredor dark pink, closed, subglobose, 1.4–3.5 × 1.5–2.5 cm; limbinside ± cream-colored with a purple zone at apex, base partlyclosed, partly convolute, margins sinuous, distal part archingand twisting, outside and inside as spathe base, or grayish, lanceolate,10–15 × 1.2–2 cm. Spadix as long as spathe; femalezone subcylindric, ca. 6 mm; ovary green or ± cream-colored,depressed obovoid, 2-ovuled; sterile zone 4–6 mm, with a fewstaminodes at base, rest grooved-ridged and set with irregular,pale purple warts; staminodes clavate, ca. 4 mm; male zone 3–5mm; appendix pink to yellow, with or without a pinkish base,slender, filiform, 5–14 cm, 0.6–3 mm in diam. at base, apexobtuse. Fl. Jun.Mountain grasslands; 1500–2700 m. S Xizang [Bangladesh, India(Sikkim), Nepal].2. Sauromatum diversifolium (Wallich ex Schott) Cusimano& Hetterscheid, Taxon 59: 445. 2010.高 原 犁 头 尖 gao yuan li tou jianTyphonium diversifolium Wallich ex Schott, Aroideae, 13.1855; Heterostalis foliolosa Schott; T. alpinum C. Y. Wu ex H.Li, Y. Xiao & X. L. Zeng; T. austrotibeticum H. Li; T. diversifoliumvar. huegelianum (Schott) Engler; T. foliolosum (Schott)Engler; T. huegelianum Schott.Underground part a subglobose tuber, 1–1.5 cm high, 0.5–1.2 cm in diam., producing annual offsets. Petiole 10–30 cm;leaf blade simple and ovate-lanceolate, 2–22 × 1–25 cm,cuneate to hastate, 3–5-lobed or pedately 5–9-sect, lobes ellipticto linear. Inflorescence appearing just before or alongsideleaves; peduncle 3–14 cm. Spathe base oblong-ovoid, 2–4 × 1–2 cm; limb outside green, sometimes flushed with purple orpurple all over, inside dark purple or greenish with dark purpleveins and flushes, oblong-lanceolate, 4–13 × 2–5 cm, apex acuminateto shortly caudate. Spadix shorter than spathe, 4–12 cm;female zone cylindric, 0.5–1.5 cm × 4–7 mm; ovary: basal halfwhite, apical half purple, ellipsoid; stigma sessile, white; sterilezone 1–3 cm, base swollen and with long staminodes, rest fusiform,with verrucate staminodal structures; proximal staminodesclavate, with only apical part distinctly and abruptlyswollen, basal part filiform, whitish with purple flushes, clavatepart yellow with a purple tip or purple; staminodal structuresconsisting of narrowly elliptic, shallow ridges with verrucatecorrugatesurface, white, pale purple, or violet; male zonecylindric, 0.6–1.3 cm × 1.5–7 mm; appendix blackish purple,broadly to narrowly cylindric, 1–7 cm × 2–8 mm, obtuse orsubacute.Open grasslands, meadows, alpine damp open ground; [ca. 2300m in Bhutan] 3300–4000 m. W Sichuan, S Xizang (Gyirong), NC Yunnan(Dayao) [Bhutan, Cambodia, NE and NW India, Myanmar, Nepal].3. Sauromatum gaoligongense Z. L. Wang & H. Li, Acta Bot.Yunnan., Suppl. 11: 61. 1999.贡 山 斑 龙 芋 gong shan ban long yuTyphonium gaoligongense (Z. L. Wang & H. Li) Hetterscheid& P. C. Boyce.Tuber depressed globose, ca. 2 cm high, 2.5–3.5 cm indiam., producing several annual offsets. Petiole uniformly greenor dark purple, to 40 cm, proximally often developing intercalary,irregular bulbils; leaf blade deeply 5–13-pedatifid;central lobe elliptic to obovate, to 12 × 5 cm; lateral lobes progressivelysmaller. Inflorescence appearing before leaves; pe-
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- Page 3 and 4: ARACEAE 523a. Female zone of spadix
- Page 5 and 6: ARACEAE 7rolled, apex rounded to ac
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- Page 15 and 16: ARACEAE 171. Calla palustris Linnae
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- Page 21 and 22: ARACEAE 23apical pore or reniform t
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- Page 25 and 26: ARACEAE 27an eccentric groove; poll
- Page 27 and 28: ARACEAE 29cal to sublateral, margin
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- Page 42 and 43: 44ARACEAE9a. Leaf blade palmate or
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- Page 47 and 48: ARACEAE 49banna) by presenting a ph
- Page 49 and 50: ARACEAE 514. Arisaema sect. Arisaem
- Page 51 and 52: ARACEAE 53Leaf solitary; petiole gr
- Page 53 and 54: ARACEAE 55slightly recurved, not au
- Page 55 and 56: ARACEAE 576. Arisaema sect. Franche
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- Page 61 and 62: ARACEAE 63Spadix unisexual; thecae
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- Page 65 and 66: ARACEAE 67stripes above neuter flow
- Page 67 and 68: ARACEAE 69narrower, with abundant a
- Page 69 and 70: ARACEAE 71Spadix up to 2.3 cm; fema
- Page 71 and 72: ARACEAE 7324. COLOCASIA Schott in S
- Page 73 and 74: ARACEAE 75sessile or subsessile, di
- Page 75 and 76: ARACEAE 77reflexed, then deliquesce
- Page 77: ARACEAE 79florescences solitary or