58ARACEAEpurple, globose, opening by an apical pore. Appendix whitishpurple proximally, dark purple distally, 6–7 cm (includingstipe), base swollen, 3–4 mm in diam., attenuate into stipe ca.10 × 1 mm. Fl. Jun–Jul.● Pinus forests, deciduous broad-leaved forests in valleys,secondary thickets; 2400–3200 m. SW Sichuan, NW Yunnan.44. Arisaema fargesii Buchet, Notul. Syst. (Paris) 1: 371. 1911.螃 蟹 七 pang xie qiArisaema bogneri P. C. Boyce & H. Li.Plants dioecious. Tuber subglobose, 3–5 cm in diam., witha few tubercles. Cataphylls 3, brown, inner one ca. 15 cm. Leafsolitary; petiole 20–40 cm, base 6–7 mm in diam., proximal 1/4sheathing into pseudostem; leaf blade 3-foliolate; leaflets sessile,membranous; central leaflet rhombic or ovate-oblong toovate, 12–32 × 9–27 cm, base shortly cuneate or combined withlateral leaflets, apex acute; lateral leaflets obliquely elliptic,outer side semiovate, 9–23 × 6–16 cm. Peduncle emerging frompseudostem, slender and shorter than petioles, 18–26 cm.Spathe purple with whitish longitudinal lines; tube subcylindric,4–8 × 1.5–2 cm, throat margins recurved and auriculate; limbarching or suberect, narrowly triangular, 6–12 × 4–4.5 cm, apexlong acuminate with a tail 1–4 cm. Spadix unisexual; femalezone ca. 2 cm; ovary obovoid; stigma subsessile, papillose;male zone cylindric, 2.5–3 cm × 4–5 mm; synandria of 2–4stamens; thecae ovoid, opening by an apical pore. Appendixrobust, suberect or slightly recurved, narrowly conic, long acuminatetoward apex, 4.5–9 cm, proximal part 7–15 mm indiam., base attenuate into a short stipe, apex obtuse, ca. 5 mmin diam. Fl. May–Jun.● Forests, thickets, in rocks; 900–2000 m. Chongqing, S Gansu,W Hubei, Hunan, Sichuan, E Xizang, NW Yunnan (Gongshan).7. Arisaema sect. Tortuosa (Engler) H. Hara, Bull. Univ. Mus. Univ. Tokyo 2: 344. 1971.曲 序 组 qu xu zuArisaema [unranked] Tortuosa Engler, Pflanzenr. 73(IV. 23F): 185. 1920; Flagellarisaema Nakai.Underground stems tuberous. Phyllotaxy quincuncial. Leaves 1 or 2(or 3); leaf blade 3-foliolate or pedate. Spathe limb erect orincurved, usually ovate. Spadix: thecae dehiscing by apical lunate slit or by pore; appendix elongate, sigmoid, tortuous, base graduallyattenuate and sessile, without neuter flowers.About 18 species: E Asia, North America; 12 species (seven endemic) in China.1a. Spathe basically purplish.2a. Flowering in autumn; spathe throat not auriculate ............................................................................................. 55. A. thunbergii2b. Flowering in spring; spathe throat broadly auriculate.3a. Spathe limb basally narrowed, not united with oblong auricles, showing 3-lobed appearance .............. 49. A. auriculatum3b. Spathe limb basally expanded and united with expanded part of spathe throat, showing singlecordate appearance ........................................................................................................................................ 56. A. cordatum1b. Spathe basically green or white.4a. Spathe white.5a. Leaf blade 3-foliolate; mature spadix bisexual ............................................................................................. 53. A. odoratum5b. Leaf blade pedately 5–7(–9)-foliolate; mature spadix female .......................................................................... 54. A. mairei4b. Spathe green.6a. Central leaflet much smaller than adjacent lateral leaflets .................................................................... 51. A. heterophyllum6b. Central leaflet ca. as large as or larger than adjacent lateral leaflets.7a. Spadix bisexual when mature ................................................................................................................ 50. A. tortuosum7b. Spadix female when mature.8a. Leaf blade (1–)3–5-foliolate; leaflets linear to narrowly oblong; ovary fusiform ............................. 45. A. saxatile8b. Leaf blade 3- or 5- or more foliolate; leaflets lanceolate to oblong; ovary bottle-shaped.9a. Spadix appendix filiform, 8–16 cm, usually pendulous.10a. Leaf blade 3-foliolate; mature spadix bisexual ................................................................... 48. A. prazeri10b. Leaf blade 5–9-foliolate; mature spadix female .................................................................. 52. A. aridum9b. Spadix appendix narrowly attenuate, less than 7 cm, curved forward.11a. Leaf blade 3-foliolate .................................................................................................... 46. A. yunnanense11b. Leaf blade 5-foliolate ............................................................................................ 47. A. quinquelobatum45. Arisaema saxatile Buchet, Notul. Syst. (Paris) 2: 124. 1911.银 南 星 yin nan xingArisaema bathycoleum Handel-Mazzetti; A. lineareBuchet.Plants dioecious. Tuber renewed seasonally, depressedglobose, 1–2 cm in diam. Cataphylls 2, creamy green, membranous,acute. Leaves 1 or 2; petiole green without spots, 9–25cm, proximally forming pseudostem; leaf blade 3-foliolate orpedately 5-foliolate; leaflets sessile or subsessile, green, narrowlylanceolate or elliptic, papery, margin entire; terminalleaflet 4–12 × 0.5–2.5 cm. Peduncle emerging from pseudostem,green, usually longer than petioles, 15–28 cm. Spathegreen or pale green; tube funnelform, 5–6 × 1–2 cm, throat mar-
ARACEAE 59gin obliquely truncate, slightly recurved; limb ovate-lanceolate,apex acute or acuminate. Spadix unisexual; female zone cylindric,2–2.5 × ca. 0.7 cm; ovary yellowish green, fusiform, styleshort; stigma capitate; male zone cylindric, ca. 2.2 cm × 4 mm;male flower of 2 or 3 stamens; anthers violet, globose, 2-thecous, dehiscing by apical pore; appendix in male spadixsessile, recurved or twining, pale green, proximally narrowlycylindric, distally filiform, to 20 cm, naked, base ca. 3 mm indiam.; appendix in female spadix recurved, usually purple, 4–5cm, much shorter than appendix in male spadix. Berries red, 1–4-seeded. Fl. Jun, fr. Aug.● Pinus forests, grassy slopes, alpine grasslands; 1600–3400 m.Sichuan, Yunnan.46. Arisaema yunnanense Buchet, Notul. Syst. (Paris) 1: 367.1911.山 珠 南 星 shan zhu nan xingArisaema talense Engler; A. talense var. latisectum Engler.Plants dioecious. Tuber renewed seasonally, pale brownoutside, white inside, depressed globose or globose, 0.5–4 cmin diam., with some offsets above; offsets sessile, globose.Cataphylls 2, green, sometimes purplish, 3–25 cm, papery, apexobtuse. Leaves 1(or 2); petiole green, without spots, 40–70 cm,basal 2/3 sheathing into pseudostem, encircled by cataphylls;leaf blade green on both surfaces, 3-foliolate; central leafletwith petiolule 0–1.5 cm, blade elliptic, sometimes ovate, 10–19× 5–12 cm, base cuneate or obtuse, apex acuminate; lateralleaflets sessile or shortly petiolulate, ovate to lanceolate, 9–15 ×5.5–9 cm, base obliquely cuneate. Peduncle emerging frompseudostem, green, without spots, longer (male) or shorter thanpetioles, 20–75 cm. Spathe green, with longitudinal white lines;tube 2–2.5 × 1–1.4 cm, throat margin obliquely truncate, notauriculate; limb arching from base, ovate, ca. 5 × 3.5 cm, apexacuminate. Spadix unisexual; female zone conic, ca. 1.5 cm;ovary green, obovoid; ovules 3, basal; style ca. 0.5 mm; stigmadepressed globose; male zone pyramidal, 1.5–2 cm, base 3–5mm in diam.; synandria of 2 or 3 stamens; thecae violet, globose,opening by an apical pore. Appendix recurved outward ordownward, pale green, cylindric, narrowed distally, 3.5–6 cm,base 2–2.5 mm in diam., naked. Berries red, subcylindric. Seeds2 or 3, ovoid-globose. Fl. May–Jul, mature fr. Aug–Sep. 2n =48.Pinus and Pinus-Quercus forests, grassy slopes, roadsides,thickets; 700–3200 m. Guizhou, Sichuan, Yunnan [N Myanmar].47. Arisaema quinquelobatum H. Li & J. Murata, sp. nov.五 叶 山 珠 南 星 wu ye shan zhu nan xingType: China. Yunnan: Lijiang, Hutiaoxia ( 虎 跳 峡 , “TigerLeaping Gorge”), 1860 m, 20 Jun 1995, J. Murata, X. Cheng,H. Takahashi, J. Ohno & H. Murata s.n. (holotype, TI).Haec species quoad inflorescentiam eadem ac Arisaemayunnanense Buchet est, sed ab eo foliis 5-foliolatis et chromosomatumnumero diploideo 2n = 24 differt.Plants dioecious, 10–40 cm tall. Tuber depressed globose,1–5 cm in diam., with some tubercles around. Leaves 1 or 2;petiole 10–45 cm, proximal 6–28 cm forming pseudostem, usuallygreenish, unspotted; leaf blade palmately 5-foliolate; leafletsgreen, narrowly elliptic to elliptic, 4–13.5 × 1–7 cm, basecuneate, margin entire, apex acuminate. Inflorescence unfoldingafter leaf blade; peduncle longer than petioles. Spathe green,with longitudinal white lines; tube 2–2.5 × 1–1.4 cm, throatmargin obliquely truncate, not auriculate; limb arching frombase, ovate, ca. 5 × 3.5 cm, apex acuminate. Spadix unisexual;female zone conic, ca. 1.5 cm; ovary green, bottle-shaped;ovules 3, basal; style ca. 0.5 mm; stigma depressed globose;male zone pale green, pyramidal, 1.5–2 cm, base 3–5 mm indiam.; synandria of 2 or 3 stamens; thecae violet or creamcolored,globose, opening by apical pore; pollen verrucate betweenconic spines. Appendix recurved outward or downward,green, narrowly conic, 3.5–6 cm, base 2–2.5 mm in diam.Berries red, subcylindric. Seeds 2 or 3, ovoid-globose. Fl. May–Jul, mature fr. Aug–Sep. 2n = 24*.● Pinus-Quercus forests, grassy slopes, roadsides, thickets; 1800–2100 m. NC and NW Yunnan (Lijiang, Wuding, Yuanmou, Zhongdian).This species is considered to be the diploid ancestor of Arisaemayunnanense, which is tetraploid, with 2n = 48 chromosomes. It alsodiffers from A. yunnanense in having always quinate leaves.48. Arisaema prazeri J. D. Hooker, Fl. Brit. India 6: 501. 1893.河 谷 南 星 he gu nan xingArisaema prazeri var. variegatum Engler; A. prazeri var.viride Engler.Plants monoecious and male. Tuber renewed seasonally,subglobose, 1–3 cm in diam. Cataphylls 2 or 3, grayish green,with dark purple striae and spots, 10–28 cm. Leaves 1 or 2;petiole grayish with purple striae and spots, 25–70 cm, proximal1/2–3/4 sheathing into pseudostem; leaf blade green, membranous,3-foliolate; leaflets subsessile; central leaflet ovatelanceolate,22–25 × 9–10 cm, base cuneate, apex acuminate;lateral leaflets obliquely ovate, 17–18 × 5–10 cm. Peduncleshorter than petioles. Spathe green with white longitudinalstripes on back; tube cylindric, 4–5 × ca. 1 cm, throat marginobliquely truncate, not auriculate; limb erect, lanceolate, 6–10 ×1.5–2.5 cm, apex acuminate, with short tail. Spadix bisexual ormale. Male spadix: male zone ca. 3 cm, base 4–5 mm in diam.;synandria of 2 stamens; thecae purple, ovoid, opening by anapical longitudinal slit. Bisexual spadix: female zone proximal,cylindric, 2.3–3 cm × 5–6 mm; ovary green, obovoid; ovules 3;stigma subsessile; male zone distal, 1–3.5 cm × ca. 4 mm;synandria as in male spadix. Appendix pendulous, usually darkpurple, filiform, to 80 cm. Fl. Jun.Tropical thickets, evergreen forests, in limestone areas; 100–1500m. Yunnan [Myanmar, Thailand].49. Arisaema auriculatum Buchet, Notul. Syst. (Paris) 2: 123.1911.长 耳 南 星 chang er nan xingArisaema auriculatum var. hungyaense (H. Li) Gusman &L. Gusman; A. hungyaense H. Li; A. omeiense P. C. Kao.Plants dioecious. Tuber globose, small, 1–2 cm in diam.,shortly stoloniferous. Cataphylls 3, brownish purple with tiny
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ARACEAE 523a. Female zone of spadix
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