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Transpiration and the Ascent

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232 7. <strong>Transpiration</strong><br />

per day. However, <strong>the</strong> actual evaporation usually is only about 80% of <strong>the</strong> <strong>the</strong>oretical<br />

maximum because evaporation also is affected by <strong>the</strong> vapor pressure gradient<br />

<strong>and</strong> by resistances in <strong>the</strong> vapor pathway. If evaporation exceeds <strong>the</strong> calculated<br />

potential rate, it often is because incident energy is supplemented by<br />

advection, <strong>the</strong> oasis effect shown in Fig. 7.2. Although <strong>the</strong> leaf area of plant<br />

st<strong>and</strong>s (leaf area index) usually is several times that of <strong>the</strong> soil on which <strong>the</strong>y<br />

grow, water loss cannot exceed that from an equal area of moist soil or a water<br />

surface receiving <strong>the</strong> same amount of energy. As <strong>the</strong> soil dries, stomatal closure<br />

reduces transpiration <strong>and</strong> drying of <strong>the</strong> soil surface reduces evaporation from<br />

<strong>the</strong> soil, hence actual ET often is considerably lower than <strong>the</strong> potential rate calculated<br />

from pan evaporation.<br />

Evaporation <strong>and</strong> transpiration losses from a field of corn <strong>and</strong> a forest are<br />

given in Tables 7.1 <strong>and</strong> 7.2, <strong>and</strong> <strong>the</strong> effect of removing plant cover in increasing<br />

runoff <strong>and</strong> evaporation is shown in Table 7.2. One might expect water losses to<br />

be similar from various types of plant cover receiving similar amounts of radiation,<br />

but variations in reflectance (albedo), roughness of <strong>the</strong> canopy, depth of<br />

rooting, <strong>and</strong> stomatal behavior result in significant differences (Rider, 1957).<br />

An extreme example is shown in Fig. 7.13. Pineapple is a CAM plant (Joshi<br />

- 3.0<br />

-28<br />

•.. -2 ..6<br />

~ -2.4<br />

E' -22<br />

E -2.0<br />

•...<br />

-E - 1..8<br />

!. -1.6<br />

o -1.4<br />

Ql<br />

~ -1.2<br />

,~ -1..0<br />

~ -08<br />

~ -06<br />

<br />

59 V'l<br />

o

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