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lichens of the lower ozark region of missouri and arkansas

lichens of the lower ozark region of missouri and arkansas

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.................................................................... A. spectabile<br />

Artho<strong>the</strong>lium spectabile A. Massal. [ARTSP] - crustose<br />

Uncommon on <strong>lower</strong> <strong>and</strong> mid boles <strong>of</strong> trees in mesic, lightly shaded areas. This species can be<br />

recognized by <strong>the</strong> well-developed, continuous, whitish gray thallus with abundant, irregularly rounded,<br />

black apo<strong>the</strong>cia typically to 0.5 × 1.5 mm.<br />

Artho<strong>the</strong>lium taediosum (Nyl.) Müll. Arg. [ARTTA] - crustose<br />

Frequent on a variety <strong>of</strong> smooth-barked trees in woodl<strong>and</strong>s, especially on Quercus coccinea, Q.<br />

rubra, <strong>and</strong> Q. velutina. This species occurs from mid boles upward into <strong>the</strong> canopy, but is almost<br />

nonexistent at <strong>the</strong> bases <strong>of</strong> trees. The apo<strong>the</strong>cia are notably smaller <strong>and</strong> more stellate-branched than<br />

in A. spectabile, with individual clusters usually < 0.5 mm broad.<br />

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ASPICILIA A. Massal. (Hymeneliaceae)<br />

Saxicolous crustose <strong>lichens</strong> with gray to grayish green, continuous to areolate thalli <strong>and</strong> usually<br />

immersed apo<strong>the</strong>cia, paraphyses moniliform, photobiont Trebouxia, asci with a slightly thickened tip,<br />

IKI-, with (4-6) 8 large, simple, hyaline, ovoid spores; 5 species in <strong>the</strong> <strong>region</strong>.<br />

1. Thallus composed <strong>of</strong> discrete, thickened, gray areoles; apo<strong>the</strong>cia immersed <strong>and</strong> sunken below level <strong>of</strong> upper<br />

cortex, pruinose, with densely pruinose thalline rim; on carbonate rocks ................. A. contorta<br />

1. Thallus continuous to rimose; apo<strong>the</strong>cia immersed but not sunken, without pruina; on siliceous rocks.<br />

2. Thallus K+ red (norstictic acid) ......................................... A. cinerea<br />

2. Thallus K- or K+ yellow (norstictic acid lacking or present as a trace substance).<br />

3. Thallus light to medium gray, K- ............................... A. caesiocinerea<br />

3. Thallus pale to dark greenish gray, K+ yellow (stictic acid).<br />

4. Thallus thin, smooth, continuous to rimose; usually near permanent water ... A. laevata<br />

4. Thallus thick, rimose to subareolate; upl<strong>and</strong>s .....................A. verrucigera<br />

Aspicilia caesiocinerea (Nyl. ex Malbr.) Arnold [ASPCAE] - crustose<br />

Occasional on shaded siliceous boulders <strong>and</strong> rock fragments in wooded upl<strong>and</strong>s, occurring on chert,<br />

granite, rhyolite, <strong>and</strong> s<strong>and</strong>stone. Interestingly, this species does not occur on massive ledges, but on<br />

smaller boulders <strong>and</strong> fragments.<br />

Aspicilia cinerea (L.) Körb. [ASPCI] - crustose<br />

Occasional, with habitats <strong>and</strong> substrates similar to those <strong>of</strong> A. caesiocinerea. This species also<br />

occurs on siliceous rocks in more moist situations along streams <strong>and</strong> seeps. [norstictic acid]<br />

Aspicilia contorta (H<strong>of</strong>fm.) Kremp. [ASPCO] - crustose<br />

Occasional on exposed dolomite boulders <strong>and</strong> ledges in glades <strong>and</strong> on bluffs, usually in full sun to<br />

light partial shade. The apo<strong>the</strong>cia are typically whi te pruinose, <strong>and</strong> overlapped around <strong>the</strong>i r margins<br />

by a zone <strong>of</strong> upper cortex that is heavily pruinose <strong>and</strong> appears almost sorediate.<br />

Aspicilia laevata (Ach.) Arnold [ASPLA] - crustose<br />

Apparently local, on hard, exposed siliceous rocks near permanent water, such as on massive igneous<br />

exposures <strong>of</strong> shut-ins along Ozark streams. The thallus <strong>of</strong> this species is greenish tinged, as contrasted<br />

with <strong>the</strong> light gray thalli <strong>of</strong> A. caesiocinerea <strong>and</strong> A. cinerea. [stictic acid, ± traces <strong>of</strong> norstictic acid]<br />

Aspicilia verrucige ra Hue [ASPVE] - crustose<br />

Rare on silicious rocks in xeric wooded upl<strong>and</strong>s. This species is similar in color <strong>and</strong> chemistry to<br />

A. laevata, but has a thicker, areolate thallus <strong>and</strong> typically occurs in upl<strong>and</strong>s, as opposed to <strong>the</strong> typical<br />

streamside habitat <strong>of</strong> A. laevata. [stictic acid, ± traces <strong>of</strong> norstictic acid]<br />

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