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north american native orchid journal - at The Culture Sheet

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Folsom: A CASE FOR EVOLUTION<br />

Casual observ<strong>at</strong>ion can often notice<br />

differences in rel<strong>at</strong>ed species. Spiranthes<br />

cernua—a compilospecies—show gene<br />

flow or influence, from the basic diploid<br />

species th<strong>at</strong> are found within its range. In<br />

the Northeast, Spiranthes cernua shows a<br />

marked influence of S. ochroleuca and in the<br />

Midwest of S. magnicamporum. Spiranthes<br />

diluvialis, an allotetraploid species derived<br />

from S. magnicamporum and S. romanzoffiana<br />

has evolved into a solid, wide-ranging<br />

species th<strong>at</strong> although still rare, often<br />

occurs now where neither ancestral parent<br />

may be found (Sheviak, 1984).<br />

Lastly, alien species have arrived<br />

whether having been borne in on storm<br />

winds or hitchhiking a ride in cargo from<br />

far off lands or brought to our shores as a<br />

garden plant. An example in Florida is<br />

Zeuxine, the lawn <strong>orchid</strong>, our only annual<br />

<strong>orchid</strong>, and has seeded in all over the place<br />

but does not drive out <strong>n<strong>at</strong>ive</strong> species.<br />

Zeuxine has been documented to have<br />

arrived in lawn seed from the Far East in<br />

the 1920s. Another example would be<br />

Oeceoclades macul<strong>at</strong>a, the African spotted<br />

<strong>orchid</strong> with its beautiful mottled leaves.<br />

This species started to appear about 25<br />

years ago and has now insinu<strong>at</strong>ed itself<br />

through central and southern Florida. It is<br />

S. diluvialis<br />

assumed it first arrived as wind-borne seed from Africa and has<br />

also colonized several islands in the Caribbean. Further <strong>north</strong><br />

Epipactis helleborine, the broad-leaved helleborine, has been known<br />

in North America for more than a century after it was first seen<br />

near Syracuse, New York. It is now found throughout most of<br />

temper<strong>at</strong>e North America and shows a decided fondness for<br />

48

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