456 THE LICHENOLOGIST Vol. 32 FIG. 9. <strong>Llimoniella</strong> ramalinae (holotypus) (in H2O). A, exciple; B, asci <strong>and</strong> paraphyses; C, ascospores. Scales: A-C= 10 (im. FIG. 10. <strong>Llimoniella</strong> ramalinae (holotypus), ascomata on thallus <strong>of</strong> Ramalina celastri. Scale = 500 urn. (Figs 9 & 10) Ascomata black, ± immersed when young, becoming sessile to shortly pedicellate, margin slightly irregular, almost disappearing when old, disc plane, becoming convex, 150-350(-600) (am diam. Exciple formed <strong>of</strong> B
2000 <strong>Skyttea</strong>, <strong>Llimoniella</strong> <strong>and</strong> Rhymbocarpus—Diederich & Etayo 457 dark, thick-walled, conglutinated hyphae, without hairs or periphysoids; lateral exciple violaceous to purplish brown, K+ green or partly violaceous, N — (brown); basal exciple greenish brown, K+ green or violaceous brown, N — (brown). Hymenium hyaline or pale violaceous, K— or partly K+ green or violaceous, c. 50 um tall, KI — . Epihymenium red-violaceous, K+ green or partly violaceous, N+ yellowish brown. Paraphyses simple or apically branched, apically swollen, without pigments. Asci clavate, wall apically not or hardly thickened, with a small but distinct ocular chamber, KI — , 50-63 x 7- 10 um, 8-spored. Ascospores hyaline, 1-septate, smooth, without a perispore, cylindrical to narrowly ellipsoid, straight or slightly curved, multiguttulate, (10-)12-16(-17) x 3-4 um. Host. Thallus <strong>of</strong> Ramalina celastri. In <strong>the</strong> recent collection, a mixture <strong>of</strong> R. celastri <strong>and</strong> R. inflata [both identified by N. Stevens], only <strong>the</strong> thalli <strong>of</strong> R. celastri are parasitized by L. ramalinae, whilst those <strong>of</strong> R. inflata are heavily attacked by a lichenicolous species <strong>of</strong> Stigmidium (possibly Arthopyrenia ramalinae Mull. Arg.). This indicates that <strong>Llimoniella</strong> ramalinae might be restricted to Ramalina celastri. Distribution. Known from two Australian localities (Victoria <strong>and</strong> Western Australia). Observations. Despite its 1-septate ascospores, this species has <strong>the</strong> same excipular structure, <strong>the</strong> same pigments <strong>and</strong> similar ascospores as <strong>the</strong> o<strong>the</strong>r species <strong>of</strong> <strong>Llimoniella</strong>, <strong>and</strong> it belongs without any doubts to this genus. In <strong>the</strong> original description, <strong>the</strong> ascomata were said to measure 500-800 um diam., but our observations show that those in <strong>the</strong> type collection are much smaller, up to 350 um only. One large ascoma <strong>of</strong> 600 um is present in <strong>the</strong> recent specimen from Victoria. Additional specimen examined. Australia: Victoria: Gunnamatta Beach, 38°23'S, 144°45'E, at sea level, s. d., Stevens 7336 (hb. Diederich). <strong>Llimoniella</strong> scabridula (Mull. Arg.) Nav.-Ros. & Haf. In Hafellner & Navarro-Rosines, Herzogia 9: 771 (1993).—Lecidea scabridula Mull. Arg., Bull. Soc. Murith. Valais 10: 55 (1881); type: Switzerl<strong>and</strong>, Valais, 'pentes gypseuses au-dessus des platreres du Granges', on Acarospora nodulosa, s. d., Wolf (G, non vid.). (Fig. 7D) For a description <strong>and</strong> additional illustrations, see Hafellner & Navarro- Rosines (1993). Hosts. Terricolous Acarospora nodulosa <strong>and</strong> A. placodiiformis (see Hafellner & Navarro-Rosines 1993). Distribution. Spain <strong>and</strong> Switzerl<strong>and</strong> (see Hafellner & Navarro-Rosines 1993).
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