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A Synopsis of the Genera Skyttea, Llimoniella and - Lichens of ...

A Synopsis of the Genera Skyttea, Llimoniella and - Lichens of ...

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466 THE LICHENOLOGIST Vol. 32<br />

Rhymbocarpus punctiformis Zopf, Hedwigia 35: 357 (1896); type: Italy, S. Tirol, St Ulrich in<br />

Groden, on Rhizocarpon geographicum, viii 1895, Arnold (M—holotypus, vid.); ibid., 10 viii 1898,<br />

Arnold Lichenes exs. 1772 (UPS—topotypus, vid.); ibid. ['Bolzano, Val Gardena (Grodner Tal),<br />

Ortisei (St Ulrich), 2-2-2 km NEE <strong>of</strong> centre, road 242 Ponte Gardena (Waidbruck)—Ortisei'], on<br />

R. geographicum, ix 1991, Triebel & Rambold 5968, 5969, 5973, 5974, 5975 (M—topotypi, vid.).<br />

(Fig. IB)<br />

For an illustration <strong>of</strong> <strong>the</strong> asci <strong>and</strong> ascospores, see Coppins et al. (1991).<br />

Ascomata aggregated in decolourized or bullate areoles <strong>of</strong> <strong>the</strong> host, at first<br />

immersed <strong>and</strong> ± peri<strong>the</strong>cioid, becoming apo<strong>the</strong>cioid when mature, normally<br />

remaining immersed, even when mature, rarely slightly erumpent, black;<br />

margin hardly visible, 50-120 (a.m diam. Exciple hyaline, except <strong>the</strong> upper part<br />

close to <strong>the</strong> epihymenium, which is greenish brown, composed <strong>of</strong> several rows<br />

<strong>of</strong> cells which are ± subglobose in <strong>the</strong> lower part <strong>and</strong> elongate in <strong>the</strong> upper<br />

part, 10-15 urn thick at <strong>the</strong> base <strong>and</strong> up to 30 \im above; excipular hairs absent<br />

or sparse, hyaline, tapered towards <strong>the</strong> apex, simple or with one septum.<br />

Subhymenium <strong>and</strong> hymenium hyaline. Epihymenium brownish to greenish.<br />

All parts KI — . Paraphyses simple to branched, hyaline, septate, slightly<br />

swollen at <strong>the</strong> apices. Asci elongate clavate, with a single functional wall<br />

layer, wall apically not thickened, I - <strong>and</strong> KI -, 8-spored. Ascospores<br />

ellipsoid, straight or slightly curved, simple, hyaline, smooth, with obtuse<br />

ends, 11-12 x 4-5 um.<br />

Hosts. Rhizocarpon atr<strong>of</strong>lavescens, R. geminatum <strong>and</strong> R. geographicum.<br />

Distribution. Greece, Italy <strong>and</strong> Sweden.<br />

Observations. The descriptions <strong>of</strong> R. geographici <strong>and</strong> R. punctiformis are<br />

extremely close. As both taxa have been collected on <strong>the</strong> same host species,<br />

<strong>the</strong>y most probably represent <strong>the</strong> same species. As <strong>the</strong> type <strong>of</strong> Nesolechia<br />

geographici is lost, we neotypify that name herewith on <strong>the</strong> type specimen <strong>of</strong><br />

Rhymbocarpus punctiformis to irrevocably fix <strong>the</strong> two names toge<strong>the</strong>r.<br />

Additional specimens examined. Sweden: Harjedalen, Tannas par., Mt Ashogen, sou<strong>the</strong>rn slope,<br />

950 m, on R. atr<strong>of</strong>lavescens, 1975, Santesson 26333 (UPS); Torne Lappmark, <strong>the</strong> Tornetrask<br />

Area, S <strong>of</strong> <strong>the</strong> railway station <strong>of</strong> Abisko, on R. geminatum, 1959, Santesson 13456a (UPS).<br />

Rhymbocarpus makarovae Diederich & Etayo sp. nov.<br />

Rhymbocarpus insignis ascomatibus immersis ad erumpentibus, atris, primo clausis, demum<br />

aperientibus, 100-200 urn, margine glabro, 3-4 pr<strong>of</strong>undis fissuris diviso ubi vetere, ascosporis<br />

ellipsoideis, aseptatis, 7-10 x 3-4 um.<br />

Typus: Russia, eastern coast <strong>of</strong> Chukchi Peninsula, Lavrentiya Bay, near Cape Krauze, on<br />

Porpidia sp., 10 July 1973, Makarova (LE—holotypus)<br />

(Fig. 13)<br />

Ascomata aggregated, immersed to erumpent, broadly attached to <strong>the</strong><br />

substratum, black, ± shiny, initially ± closed, 100-200 p.m diam.; margin,<br />

when old, divided by 3-4 deep cruciately arranged fissures, without excipular

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