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acts 24.2(14.6–37.0) cm long, narrow, elongate, ribbed, scarcely brown tomentose, without spines (rarely<br />
with few spines); rachillae 5(3–7), glabrous or scarcely tomentose initially; proximal rachillae 5.3(2.0–9.0)<br />
cm long, 0.8(0.5–1.5) mm wide; stamens 6; fruits 10.9(8.9–15.7) mm long, 7.1(5.5–10.9) mm wide, fruit<br />
surfaces uneven with numerous, subepidermal, short, often branching (Y-shaped) fibers; fruiting corollas<br />
splitting irregularly into 3 lobes, the lobes often splitting again; endocarps narrowly ellipsoid with rounded<br />
apices, the pores lateral.<br />
Subspecific variation:—Specimens occur in the western and central Amazon region in Colombia,<br />
Ecuador, Peru, Bolivia, and Brazil. Based on geography, as well as sheaths, number of pinnae, and cirri<br />
development, it is possible to recognize four subgroups—one from subAndean regions of southern Peru and<br />
Bolivia having finely and densely spiny sheaths, numerous, linear or lanceolate pinnae, and well-developed<br />
cirri; the second from the western Amazon region, mostly from the southwestern Amazon region of Brazil but<br />
also in adjacent Peru and Bolivia having non- or scarcely spiny sheaths, few, ovate pinnae, and poorlydeveloped<br />
cirri; the third from the western Amazon region of Peru and in adjacent Colombia and Brazil<br />
having non-spiny sheaths, few, ovate pinnae, and usually well-developed cirri; and the fourth from the<br />
western Amazon region in Colombia, Ecuador, and Peru, having non-spiny sheaths, numerous, lanceolate<br />
pinnae, and well-developed cirri. ANOVA shows that for pair wise comparison probabilities, six variables<br />
(plant height, petiole length, rachis length, rachis width, basal pinna length, basal pinna width) differ<br />
significantly (P