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tomentose margins (rarely without spines) (2); peduncular bracts narrow, elongate, ribbed, scarcely brown tomentose,<br />
without spines (rarely with few spines) (3); peduncular bracts broad, ridged, densely covered with short, straight,<br />
swollen-based, vertically oriented spines, these terete, whitish-brown proximally, brown distally, without tomentum (4);<br />
peduncular bracts broad, ribbed, densely brown tomentose, without spines (very rarely with a few spines) (5); peduncular<br />
bracts broad, ribbed or ridged, densely covered with felty, reddish-brown tomentum, sparsely covered with short,<br />
scarcely swollen-based, diagonally oriented, flattened spines, whitish-brown proximally, brown distally, with tomentose<br />
margins (6); peduncular bracts broad, ribbed with several more prominent, lighter colored ribs, brown, scarcely or not<br />
tomentose, without spines (rarely with a few spines) (7); peduncular bracts narrow, ribbed, densely whitish-brown<br />
tomentose, not spiny (8); peduncular bracts broad, the surfaces ribbed or ridged, brown tomentose or glabrous, sparsely<br />
to moderately covered with short, straight or sinuous, briefly swollen-based, diagonally or vertically oriented spines,<br />
these flattened or triangular in cross-section, whitish-brown proximally, black or brown distally, with tomentose margins<br />
(9); peduncular bracts broad, the surfaces deeply ridged, dark brown tomentose, sparsely covered with long, straight or<br />
sinuous spines, the bases scarcely swollen but running directly into the ridges of the bract and lying flat against the bract<br />
surface, flattened in cross-section, brown proximally and distally, with tomentose margins (10). (peduncbract)<br />
10. Rachillae brown tomentose initially (1); rachillae glabrous or scarcely tomentose initially (2). Recorded only from<br />
inflorescences with flowers because tomentum is early deciduous. (tomentum)<br />
11. Stamens five to seven (1); stamens eight to 12 (2). (stamenno)<br />
12. Fruit surfaces uneven with numerous, subepidermal, short, often branching (Y-shaped) fibers (1); fruit surfaces<br />
smooth, without any apparent subepidermal fibers (2); fruit surfaces uneven with numerous, subepidermal, long,<br />
branching fibers (3); fruit surfaces bumpy from numerous, subepidermal, short, oblique fibers (4). (fruitsurf)<br />
13. Fruiting corollas less than one quarter as long as fruits (1); fruiting corollas to half as long as fruits (2). (frcorolla)<br />
14. Fruiting corollas splitting irregularly into 3 lobes, the lobes often splitting again (1); fruiting corollas not or scarcely<br />
splitting, tending to remain cupular (2). (splitting)<br />
15. Endocarps globose to obovoid with rounded or slightly peaked apices, the pores lateral (1); endocarps narrowly<br />
ellipsoid with rounded apices, the pores lateral (2); endocarps broadly obovoid with flattened apices, the pores lateral on<br />
or near flattened apices (3); endocarps ovoid to obovoid with prominent, peaked apices, the pores lateral (4). (endocarps)<br />
Traits<br />
Abbreviations in parentheses at the end of each variable are the column labels in the Data Matrix.<br />
1. Stems solitary (1); stems clustered (2). (stems)<br />
A REVISION OF DESMONCUS (ARECACEAE)<br />
Phytotaxa 35 © 2011 <strong>Magnolia</strong> <strong>Press</strong> 51